Psephophorus | |
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Skull (LACM 162552) of P. sp. at the Natural History Museum of Los Angeles County | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | Testudines |
Suborder: | Cryptodira |
Superfamily: | Chelonioidea |
Family: | Dermochelyidae |
Genus: | † Psephophorus von Meyer, 1847 |
Type species | |
†P. polygonus | |
Species | |
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Psephophorus is an extinct genus of sea turtle that lived from the Oligocene to the Pliocene. Its remains have been found in Europe, Africa, North America, and New Zealand. [1] It was first named by Hermann von Meyer in 1847, and contains seven species, P. polygonus, P. calvertensis, P. eocaenus, P. oregonesis, [2] P. californiensis, [3] P. rupeliensis, [4] P. scaldii, [4] and a species discovered in 1995, [1] [5] P. terrypratchetti.
Psephophorus is the only Miocene dermochelyid turtle found in Europe. [6] One species of Psephophorus could measure up to ten feet in length. [7]
Von Meyer originally named Psephophorus in 1846. [8] At first he was unable to identify the creature beyond its dermal plates, but when he later received a drawing he was able to describe the specimen, which was then in Pressburg, as a fragment of a carapace, which contained seventy bones. [8]
In 1879, H. G. Seeley was asked to study the Psephophorus specimen by Franz Ritter von Hauer, the Director of the Austro-Hungarian Imperial and Royal Geological Survey. [8] Up until then, the specimen's identity had been undeterminable, with even Seeley describing it at first to seem like "the dermal covering of an Edentate closely allied to the Armadilloes." [8] Seeley examined some bone fragments and concluded the specimen was that of a reptilian creature, [8] furthermore a chelonid. It also proved to be more closely related to Sphargis than any other type in the Chelonian order. [8]
For a long time, modern Leatherback sea turtles (Dermochelys) were believed to be descended directly from Psephophorus, [9] specifically the species P. polygonus. [10] However, a 1996 analysis by Wood et al. proved that most of the taxa in the two genera were not connected, meaning Psephophorus could not be a direct ancestor of the modern leatherbacks. [9] The platelets on Psephophorus are quite similar to those on Dermochelys , despite differences in outer morphology and size. [11] The platelet comprises an external compact layer and an internal zone of cancellous bone. [11]
Sea turtles, sometimes called marine turtles, are reptiles of the order Testudines and of the suborder Cryptodira. The seven existing species of sea turtles are the flatback, green, hawksbill, leatherback, loggerhead, Kemp's ridley, and olive ridley. Six of the seven sea turtle species, all but the flatback, are present in U.S. waters, and are listed as endangered and/or threatened under the Endangered Species Act. All but the flatback turtle are listed as threatened with extinction globally on the IUCN Red List of Threatened Species. The flatback turtle is found only in the waters of Australia, Papua New Guinea, and Indonesia.
The leatherback sea turtle, sometimes called the lute turtle, leathery turtle or simply the luth, is the largest of all living turtles and the heaviest non-crocodilian reptile, reaching lengths of up to 2.7 metres and weights of 500 kilograms (1,100 lb). It is the only living species in the genus Dermochelys and family Dermochelyidae. It can easily be differentiated from other modern sea turtles by its lack of a bony shell; instead, its carapace is covered by oily flesh and flexible, leather-like skin, for which it is named. Leatherback turtles have a global range, although there are multiple distinct subpopulations. The species as a whole is considered vulnerable, and some of its subpopulations are critically endangered.
Chelidae is one of three living families of the turtle suborder Pleurodira, and are commonly called Austro-South American side-neck turtles. The family is distributed in Australia, New Guinea, parts of Indonesia, and throughout most of South America. It is a large family of turtles with a significant fossil history dating back to the Cretaceous. The family is entirely Gondwanan in origin, with no members found outside Gondwana, either in the present day or as a fossil.
Archelon is an extinct marine turtle from the Late Cretaceous, and is the largest turtle ever to have been documented, with the biggest specimen measuring 4.6 m (15 ft) from head to tail and 2.2–3.2 t in body mass. It is known only from the Pierre Shale and has one species, A. ischyros. In the past, the genus also contained A. marshii and A. copei, though these have been reassigned to Protostega and Kansastega, respectively. The genus was named in 1895 by American paleontologist George Reber Wieland based on a skeleton from South Dakota, who placed it into the extinct family Protostegidae. The leatherback sea turtle was once thought to be its closest living relative, but now, Protostegidae is thought to be a completely separate lineage from any living sea turtle.
Dermochelyidae is a family of sea turtles which has seven extinct genera and one extant genus, containing one living species, the leatherback sea turtle. The oldest fossils of the group date to the Late Cretaceous.
Macrochelys is a genus of very large freshwater turtles in the family Chelydridae, native to the Southeastern and Midwestern United States. Only a single extant species was recognized until 2014, when a study divided it into two or possibly three species. These turtles are easily recognized by three distinct dorsal ridges with raised spikes.
Protostega is an extinct genus of sea turtle containing a single species, Protostega gigas. Its fossil remains have been found in the Smoky Hill Chalk formation of western Kansas, time-equivalent beds of the Mooreville Chalk Formation of Alabama and Campanian beds of the Rybushka Formation. Fossil specimens of this species were first collected in 1871, and named by Edward Drinker Cope in 1872. With a total length of 3.9 metres (13 ft), it is the second-largest sea turtle that ever lived, second only to the giant Archelon, and one of the three largest turtle of all time along Archelon and Gigantatypus.
Protostegidae is a family of extinct marine turtles that lived during the Cretaceous period. The family includes some of the largest sea turtles that ever existed. The largest, Archelon, had a head one metre (39 in) long. Like most sea turtles, they had flattened bodies and flippers for front appendages; protostegids had minimal shells like leatherback turtles of modern times.
Santanachelys gaffneyi is an extinct species of sea turtle. It is the only species in the genus Santanachelys, which itself is a member of the extinct family Protostegidae. The species was first described from a 20-centimeter long fossil specimen unearthed in 1998 from the Santana Formation of eastern Brazil. From the rock layer from which it was excavated, it was determined that the specimen was from the Early Cretaceous period. It is therefore one of the oldest known sea turtles. but a new fossil named Desmatochelyspadillai in 2015 is estimated to be as old as 120 million years.
Zygophyseter varolai is an extinct sperm whale that lived during the Tortonian age of the Late Miocene 11.2 to 7.6 million years ago. It is known from a single specimen from the Pietra Leccese Formation in Italy. It was a member of a stem group of fossil macroraptorial sperm whales also including Brygmophyseter, Acrophyseter, and Livyatan. It probably grew to be around 6.5 to 7 meters in length and shared some characteristics with other raptorials, such as large teeth with tooth enamel that were functional in both the upper and lower jaws which the modern sperm whale lacks. It also had a beak, the ability to echolocate prey, and could have probably swum faster than the modern-day sperm whale which can reach 4 kilometers per hour (2.5 mph). These were probably used in the capture of large prey, such as large fish, seals, and whales. In fact, its common name, the killer sperm whale, refers to its feeding habits that would have had a resemblance to the modern-day killer whale.
Hesperotestudo is an extinct genus of tortoise native to North and Central America from the Early Miocene to the Late Pleistocene. Species of Hesperotestudo varied widely in size, with a large undescribed specimen from the Late Pleistocene of El Salvador reaching
Mesodermochelys is an extinct genus of sea turtle known from the Campanian to the Maastrichtian of what today is Japan and from the Maastrichtian of Chile. One species is known, the type species M. undulatus; it was given its binomial name by Ren Hirayama and Tsutomu Chitoku in 1996. Studies of its skull indicate that it was a primitive member of the Dermochelyidae that was closely related to the Protostegidae. It has been described as the best representative of Mesozoic dermochelyids.
The turtle shell is a shield for the ventral and dorsal parts of turtles, completely enclosing all the vital organs of the turtle and in some cases even the head. It is constructed of modified bony elements such as the ribs, parts of the pelvis and other bones found in most reptiles. The bone of the shell consists of both skeletal and dermal bone, showing that the complete enclosure of the shell likely evolved by including dermal armor into the rib cage.
Eunotosaurus is an extinct genus of amniote, possibly a close relative of turtles. Eunotosaurus lived in the late Middle Permian and fossils can be found in the Karoo Supergroup of South Africa and Malawi. Eunotosaurus resided in the swamps of what is now southern Africa. Its ribs were wide and flat, forming broad plates similar to a primitive turtle shell, and the vertebrae were nearly identical to those of some turtles. Accordingly, it is often considered as a possible transitional fossil between turtles and their prehistoric ancestors. However, it is possible that these turtle-like features evolved independently of the same features in turtles, since other anatomical studies and phylogenetic analyses suggest that Eunotosaurus may instead have been a parareptile, an early-diverging neodiapsid unrelated to turtles, or a synapsid.
Mesoclemmys is a genus of South American turtles in the family Chelidae.
The extant (living) reptiles of New Zealand consist of numerous species of terrestrial lizards and the lizard-like tuatara, and several species of sea turtles and sea snakes. All but one species are native to New Zealand, and all but one of the terrestrial species are endemic to New Zealand, that is, they are not found in any other country. Many species of foreign reptiles enter New Zealand through accidental importation or smuggling, but only the Australian native Lampropholis delicata has become established in the wild, and it is considered an invasive pest.
Megalochelys is an extinct genus of tortoises that lived from the Miocene to Pleistocene. They are noted for their giant size, the largest known for any tortoise, with a maximum carapace length of over 2 m (6.5 ft) in M. atlas. The genus ranged from western India and Pakistan to as far east as Sulawesi and Timor in Indonesia, though the island specimens likely represent distinct species.
The Gran Canaria giant tortoise is an extinct species of cryptodire turtle in the family Testudinidae endemic to the island of Gran Canaria, in the Canary Islands.
Erquelinnesia is an extinct turtle genus from the Cretaceous period, which has been found in fossils. Paleontologist Louis Dollo first described this genus late 19th century from fossilized samples found in the layers of Upper Paleocene / Lower Eocene in Hainaut, Belgium. Erquelinnesia lived between the Upper Cretaceous and Lower Paleocene. Louis Dollo named his discovery as Pachyrhynchus gosseleti in 1886. However, in the following year, the nomenclature was changed by Dollo to Erquelinnesia gosseleti to resolve a conflict with another already existing generic name. Moreover, an additional species was attributed to the genus Erquelinnesia. The North American species called Erquelinnesia molaria that was known only from a fossilized mandible bone has subsequently re-categorized to the genus Euclastes.
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