Archelon is an extinct marine turtle from the Late Cretaceous, and is the largest turtle ever to have been documented, with the biggest specimen measuring 4.6 m (15 ft) from head to tail and 2.2–3.2 t in body mass. It is known only from the Pierre Shale and has one species, A. ischyros. In the past, the genus also contained A. marshii and A. copei, though these have been reassigned to Protostega and Kansastega, respectively. The genus was named in 1895 by American paleontologist George Reber Wieland based on a skeleton from South Dakota, who placed it into the extinct family Protostegidae. The leatherback sea turtle was once thought to be its closest living relative, but now, Protostegidae is thought to be a completely separate lineage from any living sea turtle.
A flipper is a broad, flattened limb adapted for aquatic locomotion. It refers to the fully webbed, swimming appendages of aquatic vertebrates that are not fish.
Dermochelyidae is a family of turtles which has seven extinct genera and one extant genus, including the largest living sea turtles.
The Cryptodira are a suborder of Testudines that includes most living tortoises and turtles. Cryptodira differ from Pleurodira in that they lower their necks and pull the heads straight back into the shells, instead of folding their necks sideways along the body under the shells' marginals. They include among their species freshwater turtles, snapping turtles, tortoises, softshell turtles, and sea turtles.
Protostega is an extinct genus of sea turtle containing a single species, Protostega gigas. Its fossil remains have been found in the Smoky Hill Chalk formation of western Kansas, time-equivalent beds of the Mooreville Chalk Formation of Alabama and Campanian beds of the Rybushka Formation. Fossil specimens of this species were first collected in 1871, and named by Edward Drinker Cope in 1872. With a total length of 3.9 metres (13 ft), it is the second-largest sea turtle that ever lived, second only to the giant Archelon, and one of the three largest turtle of all time along Archelon and Gigantatypus.
Allopleuron is a genus of extinct sea turtle, which measured 2-to-2.5-metre long in life. The type species is Allopleuron hofmanni. It is a basal member of the clade Pancheloniidae, closely related to Protosphargis. Similar to Protosphargis, it was characterized by shell reduction.
Protostegidae is a family of extinct marine turtles that lived during the Cretaceous period. The family includes some of the largest sea turtles that ever existed. The largest, Archelon, had a head one metre (39 in) long. Like most sea turtles, they had flattened bodies and flippers for front appendages; protostegids had minimal shells like leatherback turtles of modern times.
Santanachelys gaffneyi is an extinct species of sea turtle. It is the only species in the genus Santanachelys, which itself is a member of the extinct family Protostegidae. The species was first described from a 20-centimeter long fossil specimen unearthed in 1998 from the Santana Formation of eastern Brazil. From the rock layer from which it was excavated, it was determined that the specimen was from the Early Cretaceous period. It is therefore one of the oldest known sea turtles. but a new fossil named Desmatochelyspadillai in 2015 is estimated to be as old as 120 million years.
Toxochelys is an extinct genus of marine turtle from the Late Cretaceous period. It is the most commonly found fossilized turtle species in the Smoky Hill Chalk, in western Kansas.
Notochelone is an extinct genus of sea turtle, which existed about 100 million years ago. The species was first described by Richard Owen in 1882 as Notochelys costata. It was renamed by Richard Lydekker in 1889. It was the most common marine reptile living in the inlands of the sea around Queensland, Australia. It was small turtle with carapace less than 1 metre (3.3 ft). Analytical studies have indicated that the creatures frequently ate benthic molluscs.
Desmatochelys is an extinct genus of sea turtles belonging to the family Protostegidae. This genus contains two known species, D. lowii and D. padillai. D. lowii was first discovered in 1895, followed by D. padillai in 2015. Having been estimated at over 120 million years old, D. padillai is currently the oldest known species of sea turtle.
Cratochelone is an extinct genus of sea turtle in the family Protostegidae and containing a single species Cratochelone berneyi. The species is known only from the mid to late Albian Toolebuc Formation, part of the Rolling Downs Group, in the Hughenden of Central northern Queensland, Australia.
Ocepechelon is an extinct genus of giant protostegid sea turtle known from Late Cretaceous phosphatic deposits of the Oulad Abdoun Basin, Khouribga Province of Morocco. It is known from the holotype OCP DEK/GE 516, a complete but isolated 70-cm-long skull, making it one of the largest marine turtles ever described. It was first named by Nathalie Bardet, Nour-Eddine Jalil, France de Lapparent de Broin, Damien Germain, Olivier Lambert and Mbarek Amaghzaz in 2013, and the type species is Ocepechelon bouyai. The feeding apparatus of Ocepechelon, a bony pipette-like snout, is unique among tetrapods and shares unique convergences with both syngnathid fishes and beaked whales.
Panchelonioidea is a clade of marine turtles that includes the sea turtles and related taxa.
Rhinochelys is an extinct genus of sea turtles belonging to the family Protostegidae.
Alienochelys is an extinct genus of sea turtle known from Maastrichian-aged Cretaceous phosphates in Morocco. With a skull measuring 41.5 centimetres (1.36 ft) long, it would have been a very large turtle. It was first described as a member of the family Dermochelyidae, meaning that it is a relative of the modern leatherback turtle, though a 2018 study identifies it as a sister taxon of Ocepechelon belonging to the family Protostegidae, indicating that it may have been closer to the extinct Archelon. Unlike other sea turtles, Alienochelys' the jaws being adapted for a powerful crushing pattern well adapted for a durophagous lifestyle, as well as its unusual cranial characteristics.
Americhelydia is a clade of turtles that consists of sea turtles, snapping turtles, the Central American river turtle and mud turtles, supported by several lines of molecular work. Prior to these studies some morphological and developmental work have considered sea turtles to be basal members of Cryptodira and kinosternids related to the trionychians in the clade Trionychoidea. Americhelydia and Testudinoidea, both clades within Durocryptodira, split a part during the early Cretaceous.
Perichelydia is a clade within Pantestudines known from the Middle Jurassic to Holocene. Alongside crown group Testudines, it also contains Helochelydridae, which is known from the Cretaceous of Europe and North America, Sichuanchelyidae from the Middle Jurassic to Paleocene of Asia and Europe, Meiolaniformes, which is known from the Cretaceous to Holocene of South America, Australia and Oceania, and Spoochelys, known from the Mid-Cretaceous Griman Creek Formation of Australia. Kallokibotion from the Late Cretaceous of Europe is also considered part of this group. Several other groups, including the proposed clade Angolachelonia, Paracryptodira, Macrobaenidae, Sinemydidae and Xinjiangchelyidae, which are sometimes considered members of Cryptodira, have also been found outside crown Testudines in several analyses. These groups are usually considered to be closer to the crown group than the other members of Perichelydia.
Thalassochelydia is a clade of extinct marine turtles from the Late Jurassic and earliest Cretaceous of Europe and South America. The group is defined as including Eurysternum, Plesiochelys and Thalassemys to the exclusion of Pelomedusa, Testudo and Protostega. While a clade uniting the families Eurysternidae, Plesiochelyidae and Thalassemydidae had been supported by phylogenetic evidence, a name was not given for the clade until 2017, when Jérémy Anquetin and colleagues coined Thalassochelydia.
Angolachelonia is a clade of extinct turtles from the Late Jurassic to Paleogene of Eurasia. The group is defined as all taxa derived from the ancestor of the type genus Angolachelys and Solnhofia, a definition that could potentially encompass a clade of entirely marine turtles. Angolachelonia was originally inclusive of only Solnhofia, Angolachelys and Sandownia when originally conceived by Octavio Mateus and colleagues in 2009, but later phylogenetic analyses by Serjocha Evers and Roger Benson in 2018 unites the family Sandownidae, including Angolachelys and Sandownia among other taxa, with the entirely Late Jurassic clade Thalassochelydia, where Solnhofia may be a basal member. While the placement of Solnhofia is weak and the clade that Angolachelonia represents may change with further analysis, the clade of Sandownidae and Thalassochelydia is well-supported, and does not collapse despite the uncertain evolutionary history of the group. Three alternative potential origins of Angolachelonia sensu Evers and Benson are shown below.
