| Gilmoremys Temporal range: Upper Cretaceous, | |
|---|---|
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Order: | Testudines |
| Suborder: | Cryptodira |
| Family: | Trionychidae |
| Subfamily: | † Plastomeninae |
| Genus: | † Gilmoremys Joyce & Lyson, 2011 |
| Species | |
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Gilmoremys is an extinct genus of softshell turtle which lived during the late Cretaceous (Maastrichtian stage) of North Dakota, Montana and Wyoming, United States.
Gilmoremys is known from five skulls, a mandible and an incomplete postcranial skeleton. The holotype of G. lancensis, USNM 6727, consists of a nearly complete carapace and an isolated hyoplastral fragment, was first assigned to the species Aspideretes lancensis. Many additional specimens were later discovered including cranial remains, and the material was assigned to its own genus, Gilmoremys. It was found from the Hell Creek Formation and from the Lance Formation (only the holotype). It was first named by Walter G. Joyce and Tyler R. Lyson in 2011 and the type species is Gilmoremys lancensis. The generic name honors Dr. Charles W. Gilmore. [1] A second species, G. gettyspherensis, is known from the late Campanian Fruitland Formation of New Mexico. [2]
Juveniles of this species have narrow skulls, a narrow processus trochlearis oticum, a deep and narrow median palatal groove, low accessory ridges, and a secondary palate fully formed by the maxilla, skeletally mature individuals have notably broad skulls, a broad processus trochlearis oticum, a shallow but broad median palatal groove, high accessory ridges, and a substantial contribution of the vomer to the secondary palate. Two large trionychid skulls and isolated shell pieces have been discovered in deposits of the Hell Creek Formation exposed in Carter County, Montana, southeastern Montana. The skulls are noticeably larger than those previously discovered and differ in their overall form by being significantly broader. Detailed analysis reveals that the new skulls and shell fragments correspond with changes in overall form representing an ontogenetic shift towards the adult morphology of this species. [3]
Cladogram after Joyce & Lyson, 2011: [1]
| Trionychidae |
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The Trionychidae are a taxonomic family of a number of turtle genera, commonly known as softshell turtles. The family was erected by Leopold Fitzinger in 1826. Softshells include some of the world's largest freshwater turtles, though many can adapt to living in highly brackish areas. Members of this family occur in Africa, Asia, and North America, with extinct species known from Australia. Most species have traditionally been included in the genus Trionyx, but the vast majority have since been moved to other genera. Among these are the North American Apalone softshells that were placed in Trionyx until 1987.
Rafetus is a genus of highly endangered softshell turtles in the family Trionychidae. It is a genus of large turtles which are found in freshwater habitats in Asia.
The Fort Union Formation is a geologic unit containing sandstones, shales, and coal beds in Wyoming, Montana, and parts of adjacent states. In the Powder River Basin, it contains important economic deposits of coal, uranium, and coalbed methane.
Emarginachelys cretacea is a turtle belonging to the group Cryptodira, known from well preserved fossils from the Maastrichtian stage of the Late Cretaceous of Montana. Its exact phylogenetic position within Cryptodira is uncertain; different authors considered it to be either the earliest described chelydrid or a fossil relative of kinosternoids.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2009.
Compsemys is an extinct genus of prehistoric turtles from the Late Cretaceous and Paleocene of North America and possibly Europe. The type species C. victa, first described by Joseph Leidy from the Hell Creek Formation in Montana in 1856, and another probable species C. russelli, described in 2012, from Paleocene deposits in France. Its affinites have long been uncertain, but it has recently been considered to be the most basal member of Paracryptodira, despite the clade first appearing in the Late Jurassic, and is sometimes included in its own family, Compsemydidae. A revision in 2020 found Compsemydidae to be more expansive, also containing Riodevemys and Selenemys from the Late Jurassic of Europe, and Peltochelys from the Early Cretaceous of Europe.
Cedrobaena is an extinct genus of turtle which existed in the Tiffanian Cedar Point Quarry, Wyoming and in the latest Maastrichtian Hell Creek Formation, United States. It was first named by Tyler R. Lyson and Walter G. Joyce in 2009 and the type species is Cedrobaena putorius.
Peckemys is an extinct genus of baenid turtle which existed in the Hell Creek Formation, United States during the late Cretaceous period. It was first named by Tyler R. Lyson and Walter G. Joyce in 2009 and the type species is Peckemys brinkman.
Gamerabaena is an extinct genus of baenid turtle which existed in North Dakota during the late Cretaceous Period. It is known from a single fragmentary skull that was found in the Maastrichtian-age Hell Creek Formation. It contains the species Gamerabaena sonsalla. Gamerabaena is similar to the genus Palatobaena, but it differs in its lack of a posterior expansion of the triturating surface, a somewhat rectangular skull, and a wide angle between the maxillae. Gamerabaena also has a lingual ridge on the inner side of the jaw that is not seen in Palatobaena.
Hutchemys is an extinct genus of softshell turtles from the late Cretaceous to the late Paleocene of New Mexico, Montana, Wyoming, Utah, and North Dakota, United States. It was first named by Walter G. Joyce, Ariel Revan, Tyler R. Lyson and Igor G. Danilov in 2009, and the type species is Hutchemys rememdium. H. rememdium is known from the holotype YPM PU 16795, which consists of a nearly complete postcranial skeleton, and from the referred specimen YPM PU 16781, found in the Ekalaka Member of the Fort Union Formation, Montana. Another referred specimen, YPM PU 14985, was found in the Cedar Point Quarry, Wyoming. The second species, H. arctochelys, is known from the holotype YPM PU 16319, a nearly complete carapace, and from the paratypes YPM PU 16320, YPM PU 16321, YPM PU 16322, YPM PU 16238. All specimens of H. arctochelys were recovered from the same quarry of the Tongue River Member, Fort Union Formation, near Burns Mine of Montana. A possible third species is represented by the unnamed specimen UCMP 130000 from the Paleocene Tullock Formation of Montana. Aspideretes? nassau (YPM PU 11566) from the Fort Union Formation, Duffy's Ranch of Sweet Grass County, Montana was also assigned to Hutchemys sp. A fourth species of Hutchemys, Hutchemys walkerorum, has been uncovered from the Hell Creek Formation of North Dakota. H. walkerorum is known from the holotype BDM 063, identified by the discovery and assembly of portions of its carapace. The fossil of H. walkerorum suggests that it was the only Hutchemys to live exclusively during the late Cretaceous era.
Palatobaena is an extinct genus of baenid turtle. It was first named by Gaffney in 1972 and the type species is Palatobaena bairdi. It based on a fragmentary skull from the Fort Union Formation of the Bighorn Basin of Wyoming. The two other species are P. gaffneyi and P. cohen which existed in Hell Creek Formation, North Dakota during the late Cretaceous period.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2011.
Cyclanorbinae, also known commonly as the flapshell turtles, is a subfamily of softshell turtles in the family Trionychidae. The subfamily is native to Africa and Asia.
The Trionychinae are a subfamily of turtles in the family Trionychidae.
Khunnuchelys was a genus of trionychine turtle from the Late Cretaceous of Asia. Three species are known, K. erinhotensis, the type species, K. kizylkumensis, and K. lophorhothon. K. erinhotensis is known from the Iren Dabasu Formation in China from the late Turonian until the middle Campanian. K. kizylkumensis is known from the late Turonian Bissekty Formation of Uzbekistan. The third species, described in 2013 by Danilov et al., is known from the early to middle Campanian aged Bostobe Formation of Kazakhstan.
Neurankylus is an extinct genus of turtles in the family Baenidae that lived between 112 and 61 million years ago in Canada and the United States. It was originally placed within the monotypic family Neurankylidae, but it has since been placed in the Neurankylinae, alongside Trinitichelys. The type species, Neurankylus eximius, was described by Lawrence Lambe in 1902. The species N. lithographicus was discovered in the Milk River Formation (Canada), alongside the holotype of the pachycephalosaurid dinosaur Acrotholus audeti.
Anatolemys is an extinct turtle genus in the family Macrobaenidae. Two species are known, both of which lived in the Late Cretaceous. Fossils were discovered in the Yalovach Formation of Tajikistan, the Kulbikin Member and Khodzhakul and Bissekty Formations of Uzbekistan and the Bostobe Formation of Kazakhstan. With 70 cm (2.3 ft) in carapace length, Anatolemys maximus was one of the three largest macrobaenids along with Early Cretaceous Yakemys multiporcata and Paleocene Judithemys backmani.
Perichelydia is a clade within Pantestudines known from the Middle Jurassic to Holocene. Alongside crown group Testudines, it also contains Helochelydridae, which is known from the Cretaceous of Europe and North America, Sichuanchelyidae from the Middle Jurassic to Paleocene of Asia and Europe, Meiolaniformes, which is known from the Cretaceous to Holocene of South America, Australia and Oceania, and Spoochelys, known from the Mid-Cretaceous Griman Creek Formation of Australia. Kallokibotion from the Late Cretaceous of Europe is also considered part of this group. Several other groups, including the proposed clade Angolachelonia, Paracryptodira, Macrobaenidae, Sinemydidae and Xinjiangchelyidae, which are sometimes considered members of Cryptodira, have also been found outside crown Testudines in several analyses. These groups are usually considered to be closer to the crown group than the other members of Perichelydia.
Drazinderetes is a large bodied genus of soft shell turtle from the Middle Eocene Drazinda Formation of Pakistan. Its presence in the shallow marine deposits of the Drazinda Formation suggests that Drazinderetes may have been a partially or fully marine animal. Indetermined trionychine remains from the same formation may suggest that Drazinderetes could have been among the largest known turtles, with one entoplastron indicating a potential length of 1.5 to 2.1 meters. Drazinderetes currently consists of only a single species: Drazinderetes tethyensis.
Plastomeninae is an extinct subfamily of softshell turtles that inhabited most of North America from the Cretaceous to the Eocene. Members of this subfamily are also known as plastomenines.
Trionychidae family | ||||
|---|---|---|---|---|
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| Genera |
|  | ||
| Amyda | ||||
| Apalone | ||||
| † Axestemys | ||||
| Chitra | ||||
| Cyclanorbis | ||||
| Cycloderma | ||||
| Dogania | ||||
| † Gilmoremys | ||||
| † Hutchemys | ||||
| † Khunnuchelys | ||||
| Lissemys | ||||
| Nilssonia | ||||
| † Palaeoamyda | ||||
| Palea | ||||
| Pelochelys | ||||
| Pelodiscus | ||||
| Rafetus | ||||
| Trionyx | ||||
Phylogenetic arrangement of turtles based on turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status. †=extinct. | ||||
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