The Trionychidae are a taxonomic family of a number of turtle genera, commonly known as softshell turtles. The family was erected by Leopold Fitzinger in 1826. Softshells include some of the world's largest freshwater turtles, though many can adapt to living in highly brackish areas. Members of this family occur in Africa, Asia, and North America, with extinct species known from Australia. Most species have traditionally been included in the genus Trionyx, but the vast majority have since been moved to other genera. Among these are the North American Apalone softshells that were placed in Trionyx until 1987.
Archelon is an extinct marine turtle from the Late Cretaceous, and is the largest turtle ever to have been documented, with the biggest specimen measuring 4.6 m (15 ft) from head to tail and 2.2–3.2 t in body mass. It is known only from the Pierre Shale and has one species, A. ischyros. In the past, the genus also contained A. marshii and A. copei, though these have been reassigned to Protostega and Kansastega, respectively. The genus was named in 1895 by American paleontologist George Reber Wieland based on a skeleton from South Dakota, who placed it into the extinct family Protostegidae. The leatherback sea turtle was once thought to be its closest living relative, but now, Protostegidae is thought to be a completely separate lineage from any living sea turtle.
The pig-nosed turtle, also known as the pitted-shelled turtle or Fly River turtle, is a species of turtle native to northern Australia and southern New Guinea. It is the only living member of the family Carettochelyidae, which are related to softshell turtles.
Leith's softshell turtle is a species of turtle in the family Trionychidae. The species is found in peninsular Indian rivers including the Thungabhadra, Ghataprabha, Bhavani, Godavari, Kaveri and Moyar Rivers. The type locality is Pune in India.
The Indian softshell turtle, or Ganges softshell turtle is a species of softshell turtle found in South Asia in rivers such as the Ganges, Indus and Mahanadi. This vulnerable turtle reaches a carapace length of up to 94 cm (37 in). It feeds mostly on fish, amphibians, carrion and other animal matter, but also takes aquatic plants. This turtle is listed in part II of Schedule I of the Wild Life (Protection) Act, 1972 and possession of this species is an offence.
Stupendemys is an extinct genus of freshwater side-necked turtle, belonging to the family Podocnemididae. It is the largest freshwater turtle known to have existed, with a carapace over 2 meters long. Its fossils have been found in northern South America, in rocks dating from the Middle Miocene to the very start of the Pliocene, about 13 to 5 million years ago. Male specimens are known to have possessed bony horns growing from the front edges of the shell and the discovery of the fossil of a young adult shows that the carapace of these turtles flattens with age. A fossil skull described in 2021 indicates that Stupendemys was a generalist feeder.
Axestemys is an extinct genus of softshell turtle that lived from the Late Cretaceous to the Eocene in western North America and Europe.
The spiny softshell turtle is a species of softshell turtle, one of the largest freshwater turtle species in North America. Both the common name, spiny softshell, and the specific name, spinifera (spine-bearing), refer to the spiny, cone-like projections on the leading edge of the carapace, which are not scutes (scales).
The Florida softshell turtle is a species of turtle in the family Trionychidae. The species is native to the Southeastern United States.
The Yangtze giant softshell turtle, also known commonly as the Red River giant softshell turtle, the Shanghai softshell turtle, the speckled softshell turtle, and Swinhoe's softshell turtle, is an extremely rare species of turtle in the family Trionychidae. It may be the largest living freshwater turtle in the world. The species is native to eastern and southern China and northern Vietnam. With a known population of only three individuals, and with the last females dying out, this species is considered functionally extinct.
Nilssonia is a genus of softshell turtles from rivers, streams, ponds, and lakes in South Asia and Burma. In many treatments, it is monotypic, with the single species Burmese peacock softshell. However, the supposed other genus of peacock softshells, Aspideretes, is more closely related to N. formosa than had been believed. They differ only in the neural plates between the first pleural scale pair of the bony carapace, which are fused into one in N. formosa and unfused in the others.
Niolamia is an extinct genus of South American meiolaniid turtle with a long and complex history. Like its relatives, Niolamia was a robust and heavily armored terrestrial turtle with large, horn like scales covering its head and a tail encased by rings of bone. This heavily armored build may have served the animal during intraspecific combat during courtship. Scans of the skull reveal that Niolamia likely had a great sense of smell but only low frequency hearing, indicating that these animals communicated more through chemical signals and smells than through sound.
The smooth softshell turtle is a species of softshell turtle of the family Trionychidae. This freshwater turtle is endemic to the United States where it inhabits the Mississippi River system and other river basins emptying into the Gulf of Mexico.
Eileanchelys is an extinct genus of primitive turtle from the Middle Jurassic (Bathonian) period some 164 million years ago of Britain. Only one species is recorded, Eileanchelys waldmani. It is the best-represented turtle from the Middle Jurassic, because of the amount of specimens that can be assigned to it. The turtle is also one of the oldest turtles ever found to be aquatic, and might represent a milestone in turtle evolution.
Hutchemys is an extinct genus of softshell turtles from the late Cretaceous to the late Paleocene of New Mexico, Montana, Wyoming, Utah, and North Dakota, United States. It was first named by Walter G. Joyce, Ariel Revan, Tyler R. Lyson and Igor G. Danilov in 2009, and the type species is Hutchemys rememdium. H. rememdium is known from the holotype YPM PU 16795, which consists of a nearly complete postcranial skeleton, and from the referred specimen YPM PU 16781, found in the Ekalaka Member of the Fort Union Formation, Montana. Another referred specimen, YPM PU 14985, was found in the Cedar Point Quarry, Wyoming. The second species, H. arctochelys, is known from the holotype YPM PU 16319, a nearly complete carapace, and from the paratypes YPM PU 16320, YPM PU 16321, YPM PU 16322, YPM PU 16238. All specimens of H. arctochelys were recovered from the same quarry of the Tongue River Member, Fort Union Formation, near Burns Mine of Montana. A possible third species is represented by the unnamed specimen UCMP 130000 from the Paleocene Tullock Formation of Montana. Aspideretes? nassau (YPM PU 11566) from the Fort Union Formation, Duffy's Ranch of Sweet Grass County, Montana was also assigned to Hutchemys sp. A fourth species of Hutchemys, Hutchemys walkerorum, has been uncovered from the Hell Creek Formation of North Dakota. H. walkerorum is known from the holotype BDM 063, identified by the discovery and assembly of portions of its carapace. The fossil of H. walkerorum suggests that it was the only Hutchemys to live exclusively during the late Cretaceous era.
The Trionychinae are a subfamily of turtles in the family Trionychidae.
Trionyx is a genus of softshell turtles belonging to the family Trionychidae. In the past many species in the family were classified in this genus, but today T. triunguis, the African or Nile softshell turtle, is the only extant softshell still classified as Trionyx. The other species still assigned to this genus are only known from fossils. T. triunguis is a relatively large, aquatic piscivore.
Gilmoremys is an extinct genus of softshell turtle which lived during the late Cretaceous of North Dakota, Montana and Wyoming, United States.
Plastomeninae is an extinct subfamily of softshell turtles that inhabited most of North America from the Cretaceous to the Eocene. Members of this subfamily are also known as plastomenines.
Leviathanochelys is an extinct genus of giant sea turtle from the Middle Campanian of northern Spain. Although only known from the rear end of the carapace and the connecting pubic bones, these elements clearly show that they belonged to a turtle of great size, comparable in dimensions to the giant Archelon and Protostega from the Western Interior Seaway of the USA. This makes Leviathanochelys the first giant sea turtle known from Europe and one of the largest known turtles of all time. This great size was evolved independently from the American protostegid turtles and was likely an adaptation to the conditions of the Cretaceous oceans covering much of Europe. Being classified in the superfamily Chelonioidea, Leviathanochelys is likely more closely related to modern sea turtles than the protostegids, which are thought to be more basal and have a more uncertain relationship to modern sea turtles. Leviathanochelys is thought to have been a pelagic animal. The genus contains a single species, Leviathanochelys aenigmatica.
