Aetonyx

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Aetonyx
Temporal range: Early Jurassic, HettangianSinemurian 201.4–192.9  Ma
Broom 1911 plate XVII.jpg
Type specimens of Aetonyx palustris (19–23), Massospondylus harriesi (18), and Geranosaurus (24)
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Eusaurischia
Clade: Sauropodomorpha
Genus: Aetonyx
Broom, 1911 [1]
Species
  • A. palustrisBroom, 1911 [1]

Aetonyx is a dubious genus of sauropodomorph dinosaur from the Early Jurassic of southern Africa. Its only species is A. palustris, which was named by Robert Broom in 1911 based on a fragmentary skeleton from the upper Elliot Formation found near Fouriesburg, Free State Province. Broom considered it as a species of "carnivorous dinosaur". In 1924, Sydney H. Haughton assigned a second specimen to Aetonyx, which is also from Fouriesburg. In 1932, Friedrich von Huene suggested that the species Thecodontosaurus dubius , which Haughton had named in his 1924 paper, is a synonym of Aetonyx palustris. The species was later synonymised with Massospondylus harriesi and Massospondylus carinatus . A 2004 review lists it as an indeterminate sauropodomorph.

Contents

Discovery

Aetonyx was named in 1911 by Robert Broom from a fragmentary skeleton excavated by A. R. Walker, who worked at the Iziko South African Museum, where the specimen is still stored (specimen number SAM-PK-2768-2770). [1] :294,304 [2] :173 The specimen was found near Fouriesburg, Free State Province, in sediments of the upper Elliot Formation, which was deposited during the Hettangian and Sinemurian ages of the Early Jurassic. [2] :173 [3] The specimen is well-preserved and consists of three neck vertebrae, one back vertebra, some distal tail vertebrae, a shoulder blade with coracoid , and parts of the forelimb ( humerus , radius , ulna , both hands) and hind limb (the upper end of a tibia and an almost complete foot). The humerus was 17.4 cm in length. [4] :91 Broom did not provide an etymology of the name, but Donald Glut, in a 1997 popular book, suggested that it derives from the Latin aetatis'old' and the Ancient Greek onyx'claw'. [5]

Taxonomic history and status

Taxa today classified as basal sauropodomorphs, including Aetonyx, have been historically classified as theropods until the mid-twentieth century. [6] :102 Broom introduced the new taxon as a "carnivorous dinosaur". [1] A second specimen from the same locality was assigned to Aetonyx in 1924 by Sydney H. Haughton; this specimen consists of the lower end of a fibula and three right metatarsals . [7] [4] :91 In 1932, Friedrich von Huene classified Aetonyx within the Carnosauria, noting resemblances to the large carnivorous theropod Megalosaurus . The penultimate phalanges of the hand are elongated in Aetonyx, which von Huene thought was an adaptation for capturing prey. He cautioned, however, that only the discovery of a skull can confirm its carnosaurian affinities. Von Huene also declared Thecodontosaurus dubius , a species that Haughton had named in his 1924 paper, to be a synonym of Aetonyx palustris. T. dubius was based on a partly articulated (connected) skeleton from the Clarens Formation found near Ladybrand, South Africa [4] :91–93 [8]

In 1970, Rodney Steel still listed Aetonyx palustris as a valid taxon, but in 1976, Peter Galton and Michael Albert Cluver proposed that it is a synonym of Massospondylus harriesi , a species that was named by Broom in 1911, in the same paper that named Aetonyx palustris itself. [9] :47 [8] :146 [1] In 1981, Michael Cooper synonymised both species with the type species of Massospondylus, M. carinatus, which was followed by Galton in a 1990 review. [10] [11] In a 2004 review, Galton and Paul Upchurch considered all these remains to be undiagnostic and listed Aetonyx palustris, Thecodontosaurus dubius, and Massospondylus harriesi as indeterminate sauropodomorphs (Nomina dubia). [12] [2] :173

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References

  1. 1 2 3 4 5 Broom, R. (1911). "On the dinosaurs of the Stormberg, South Africa" (PDF). Annals of the South African Museum. 7: 291–308.
  2. 1 2 3 Barrett, Paul M.; Chapelle, Kimberley EJ; Staunton, Casey K.; Botha, Jennifer; Choiniere, Jonah N. (2019). "Postcranial osteology of the neotype specimen of Massospondylus carinatus Owen, 1854 (Dinosauria: Sauropodomorpha) from the upper Elliot formation of South Africa". Palaeontologia Africana. 53: 174.
  3. Bordy, Emese M.; Smith, Roger M. H.; Choiniere, Jonah N.; Rubidge, Bruce S. (2024). "Selected Karoo geoheritage sites of palaeontological significance in South Africa and Lesotho". Geological Society, London, Special Publications. 543 (1): 431–446. doi:10.1144/SP543-2022-202.
  4. 1 2 3 Huene, F. von (1932). Wolfgang Soergel (ed.). "Die fossile Reptil-Ordnung Saurischia, ihre Entwicklung und Geschichte" [The fossil reptile order Saurischia, its evolution and history]. Monographien zur Geologie und Palaeontologie (in German). Ser. 1 (4): 123–124.
  5. Glut, Donald F. (1997). "Aetonyx". Dinosaurs: The Encyclopedia. Jefferson, North Carolina: McFarland & Co. p. 87. ISBN   978-0-89950-917-4.
  6. Barrett, Paul M.; Chapelle, Kimberley EJ (2024). "A brief history of Massospondylus: its discovery, historical taxonomy and redescription of the original syntype series". Palaeontologia africana. 58: 97–131. hdl:10539/43016.
  7. Haughton, Sydney H. (1924). "The fauna and stratigraphy of the Stormberg Series". Annals of the South African Museum. 12: 323–497.
  8. 1 2 Galton, Peter M.; Cluver, Michael Albert (1976). "Anchisaurus capensis (Broom) and a revision of the Anchisauridae (Reptilia, Saurischia)". Annals of the South African Museum. 69 (6): 121–159.
  9. Steel, R. (1970). "Part 14. Saurischia". Handbuch der Paläoherpetologie/Encyclopedia of Paleoherpetology. Stuttgart: Gustav Fischer Verlag. pp. 1–87.
  10. Cooper, M. R. (1981). "The prosauropod dinosaur Massospondylus carinatus Owen from Zimbabwe: its biology, mode of life and phylogenetic significance". Occasional Papers of the National Museums and Monuments of Rhodesia, Series B, Natural Sciences. 6 (10): 689–840.
  11. Galton, Peter M. (1990). "Basal Sauropodomorpha-Prosauropoda". In Weishampel, D.B.; Dodson, P.; Osmólska, H. (eds.). The Dinosauria (1 ed.). Berkeley: University of California Press. pp. 320–344. ISBN   978-0-520-06727-1.
  12. Galton, Peter M.; Upchurch, Paul (2004). "Prosauropoda". In Weishampel, D.B.; Dodson, P.; Osmólska, H. (eds.). The Dinosauria (2 ed.). Berkeley: University of California Press. pp. 232–258. ISBN   978-0-520-25408-4.