Hyoliths are animals with small conical shells, known as fossils from the Palaeozoic Era. They are lophophorates, a group which includes the brachiopods.
A sclerite is a hardened body part. In various branches of biology the term is applied to various structures, but not as a rule to vertebrate anatomical features such as bones and teeth. Instead it refers most commonly to the hardened parts of arthropod exoskeletons and the internal spicules of invertebrates such as certain sponges and soft corals. In paleontology, a scleritome is the complete set of sclerites of an organism, often all that is known from fossil invertebrates.
Microdictyon is an extinct "armored worm" coated with net-like scleritic scales, known from the Early Cambrian Maotianshan shale of Yunnan China and other parts of the world. Microdictyon is sometimes included in a somewhat ill-defined Phylum – Lobopodia – that includes several other odd worm-like and segmented free-swimming animals that do not appear to be arthropods or worms. The phylum includes Microdictyon, Onychodictyon, Cardiodictyon, Luolishania, and Paucipodia. The isolated sclerites of Microdictyon are known from other Lower Cambrian deposits. Microdictyon sclerites appear to have moulted; one sclerite seems to have been preserved during ecdysis.
The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is Halkieria, which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the small shelly fossil assemblages. The best known species is Halkieria evangelista, from the North Greenland Sirius Passet Lagerstätte, in which complete specimens were collected on an expedition in 1989. The fossils were described by Simon Conway Morris and John Peel in a short paper in 1990 in the journal Nature. Later a more thorough description was undertaken in 1995 in the journal Philosophical Transactions of the Royal Society of London and wider evolutionary implications were posed.
Halwaxiida or halwaxiids is a proposed clade equivalent to the older orders Sachitida He 1980 and Thambetolepidea Jell 1981, loosely uniting scale-bearing Cambrian animals, which may lie in the stem group to molluscs or lophotrochozoa. Some palaeontologists question the validity of the Halwaxiida clade.
The Chancelloriids are an extinct family of animal common in sediments from the Early Cambrian to the early Late Cambrian. Many of these fossils consists only of spines and other fragments, and it is not certain that they belong to the same type of organism. Other specimens appear to be more complete and to represent sessile, bag-like organisms with a soft skin armored with star-shaped calcareous sclerites from which radiate sharp spines.
Shelly limestone is a highly fossiliferous limestone, composed of a number of fossilized organisms such as brachiopods, bryozoans, crinoids, sponges, corals and mollusks. It varies in color, texture and hardness. Coquina is a poorly indurated form of shelly limestone.
The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Cambrian Period. They are very diverse, and there is no formal definition of "small shelly fauna" or "small shelly fossils". Almost all are from earlier rocks than more familiar fossils such as trilobites. Since most SSFs were preserved by being covered quickly with phosphate and this method of preservation is mainly limited to the Late Ediacaran and Early Cambrian periods, the animals that made them may actually have arisen earlier and persisted after this time span.
Volborthella is an animal of uncertain classification, whose fossils pre-date 530 million years ago. It has been considered for a period a cephalopod. However discoveries of more detailed fossils showed that Volborthella’s small, conical shell was not secreted but built from grains of the mineral silicon dioxide (silica), and that it was not divided into a series of compartments by septa as those of fossil shelled cephalopods and the living Nautilus are. This illusion was a result of the laminated texture of the organisms' tests. Therefore, Volborthella’s classification is now uncertain. It has been speculated that it may in fact represent a sclerite of a larger organism, on the basis of one specimen; however, it may be premature to accept this hypothesis, as the arrangement of sclerites producing this impression may have occurred by chance. The Ordovician scleritome-bearing Curviconophorus, as well as the Halwaxiids, lobopods and echinoderms, demonstrate the diversity of organisms which may produce a scleritome of this nature. The related Campitius was originally suggested to be part of a radula rather than a scleritome, but is now considered a synonym of Volborthella.
Tommotiids are Cambrian (Terreneuvian) shelly fossils thought to belong to the Brachiopod + Phoronid lineage (Brachiozoa).
The palaeoscolecids are a group of extinct ecdysozoan worms resembling armoured priapulids. They are known from the Lower Cambrian to the late Silurian; they are mainly found as disarticulated sclerites, but are also preserved in many of the Cambrian lagerstätten. They take their name from the typifying genus Palaeoscolex.
The Mount Cap Formation is a geologic formation exposed in the Mackenzie Mountains, northern Canada. It was deposited in a shallow shelf setting in the late Early Cambrian, and contains an array of Burgess Shale-type microfossils that have been recovered by acid maceration.
Sachites is an extinct genus of halkeriid that is only known from fossilised spiny sclerites; many Sachites specimens are now referred to as other halkieriid taxa. Although believed to be related to the halkieriids, a chancelloriid affinity has more recently been proposed.
Mongolitubulus is a form genus encapsulating a range of ornamented conical small shelly fossils of the Cambrian period. It is potentially synonymous with Rushtonites, Tubuterium and certain species of Rhombocorniculum, and owing to the similarity of the genera, they are all dealt with herein. Organisms that bore Mongolitubulus-like projections include trilobites, bradoriid arthropods and hallucigeniid lobopodians.
Mickwitziids are a Cambrian group of shelly fossils with originally phosphatic valves, belonging to the Brachiopod stem group, and exemplified by the genus Mickwitzia – the other genera are Heliomedusa and Setatella. The family Mickwitziidae is conceivably paraphyletic with respect to certain crown-group brachiopods.
Rhombocorniculum is a species of small shelly fossil comprising twisted ornamented cones. It has been described from the Comely limestone and elsewhere. R. cancellatum straddles the Atdabanian/Botomian boundary. The structure of its inner layer suggests that its phosphatic fibres formed within a flexible organic matrix.
Tannuolina is a genus of tommotiid, belonging to the brachiopod stem lineage.
The camenellans, consisting of the genera Camenalla, Dailyatia, Kennardia, Kelanella and Lapworthella, are a group of Tommotiid invertebrates from the Cambrian period, reconstructed as sister to all others. They are known from isolated sclerites, but are believed to have a scleritomous, Halkieria-like construction.
Nidelric is a Cambrian genus of scleritomous organism, tentatively interpreted as a chancelloriid.
