Virginia chain fern | |
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Sori | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Division: | Polypodiophyta |
Class: | Polypodiopsida |
Order: | Polypodiales |
Suborder: | Aspleniineae |
Family: | Blechnaceae |
Genus: | Anchistea C.Presl [2] |
Species: | A. virginica |
Binomial name | |
Anchistea virginica (L.) C.Presl [2] | |
Synonyms [2] [3] | |
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Anchistea is a genus of leptosporangiate ferns in the family Blechnaceae. It has only one species, Anchistea virginica (synonym Woodwardia virginica) the Virginia chain fern, which has long creeping, scaly, underground stems or rhizomes giving rise to tall (up to about 4 feet, 120 centimetres) widely separated, deciduous, single leaves. In contrast, the leaves of Osmundastrum cinnamomeum , which can be mistaken for A. virginica, grow in a group from a crown. Also in contrast to O. cinnamomeum the leaves are monomorphic without distinct fertile fronds. The lower petiole or stipe is dark purple to black, shiny and swollen, the upper rachis is dull green. The leaf blade is green and lanceolate, composed of 12 to 23 paired, alternate pinnatifid pinnae. The pinnae are subdivided into 15 to 20 paired segments that are ovate to oblong. The lower rachis is naked for about half its length. The sori or spore-producing bodies are found on the underside of the pinnae and are long and form a double row which outlines the major veins of the pinnae. In common with all ferns, A. virginica exhibits a gametophyte stage in its life cycle (alternation of generations) and develops a haploid reproductive prothallus as an independent plant. The spores are produced in red-brown sori which line the spaces (areolae) between the costa and costules. Further photographs can be found at the Connecticut Botanical Society and Ontario Ferns websites.
Anchistea virginica was first described by Carl Linnaeus in 1771 as Blechnum virginicum. [4] It was transferred to Woodwardia by James Edward Smith in 1793, [5] and to Anchistea by Carl Presl in 1851. [2] [6] Some sources retain it in Woodwardia, [3] but the Pteridophyte Phylogeny Group classification of 2016 (PPG I) retains Presl's genus, making it the sole species in Anchistea. [2]
Fossils of Anchistea virginica have been described from the Middle Miocene of the U.S. states of Washington and Mississippi, suggesting its range was probably once much more widespread in North America. [7] [8] In the Middle Miocene, Anchistea virginica apparently occupied similar habitats as in the present day.
Endemic to eastern North America from Florida to Nova Scotia and west to Michigan and Illinois. A. virginica is mostly found on the Atlantic coastal plain and Piedmont although it also grows in the eastern Great Lakes region as far west as Illinois. The Flora of North America website has a distribution map.
Anchistea virginica grows in the wet soils of open wet swampy woods, acid bogs, and along streams and roadside ditches, [9] avoiding calcareous substrates. A. virginica is an important constituent of the field layer of flatwoods, Atlantic white cedar ( Chamaecyparis thyoides ) swamp forest and bay ( Magnolia virginiana ) forests. [10]
The plant is occasionally cultivated as a greenhouse or garden ornamental. Hardy to USDA Zone 3.
Asplenium platyneuron, commonly known as ebony spleenwort or brownstem spleenwort, is a fern native to North America east of the Rocky Mountains. It takes its common name from its dark, reddish-brown, glossy stipe and rachis, which support a once-divided, pinnate leaf. The fertile fronds, which die off in the winter, are darker green and stand upright, while the sterile fronds are evergreen and lie flat on the ground. An auricle at the base of each pinna points towards the tip of the frond. The dimorphic fronds and alternate, rather than opposite, pinnae distinguish it from the similar black-stemmed spleenwort.
Blechnaceae is a family of ferns in the order Polypodiales, with a cosmopolitan distribution. Its status as a family and the number of genera included have both varied considerably. In the Pteridophyte Phylogeny Group classification of 2016, the family has 24 genera, and excludes genera placed in the separate family Onocleaceae. The family is divided into three subfamilies, including Blechnoideae s.s. Alternatively, the entire family may be treated as the subfamily Blechnoideae s.l. of a very broadly defined family Aspleniaceae, and include genera others place in Onocleaceae.
Dryopteris filix-mas, the male fern, is a common fern of the temperate Northern Hemisphere, native to much of Europe, Asia, and North America. It favours damp shaded areas in the understory of woodlands, but also shady places on hedge-banks, and rocks, and screes. Near the northern limit of its distribution it prefers sunny, well-drained sites. It is much less abundant in North America than in Europe.
Osmundastrum is genus of leptosporangiate ferns in the family Osmundaceae with one living species, Osmundastrum cinnamomeum, the cinnamon fern. It is native to the Americas and eastern Asia, growing in swamps, bogs and moist woodlands.
Frond dimorphism refers to a difference in ferns between the fertile and sterile fronds. Since ferns, unlike flowering plants, bear spores on the leaf blade itself, this may affect the form of the frond itself. In some species of ferns, there is virtually no difference between the fertile and sterile fronds, such as in the genus Dryopteris, other than the mere presence of the sori, or fruit-dots, on the back of the fronds. Some other species, such as Polystichum acrostichoides, or some ferns of the genus Osmunda, feature dimorphism on a portion of the frond only. Others, such as some species of Blechnum and Woodwardia, have fertile fronds that are markedly taller than the sterile. Still others, such as Osmunda cinnamomea, or plants of the family Onocleaceae, have fertile fronds that are completely different from the sterile.
Woodwardia is a genus of ferns in the family Blechnaceae, in the suborder Aspleniineae of the order Polypodiales. Species are known as netted-chain ferns. The genus is native to warm temperate and subtropical regions of the Northern Hemisphere. They are large ferns, with fronds growing to 50–300 cm long depending on the species. The fossil record of the genus extends to the Paleocene.
Blechnum, known as hard fern, is a genus of ferns in the family Blechnaceae, subfamily Blechnoideae, according to the Pteridophyte Phylogeny Group classification of 2016. Two very different circumscriptions of the genus are used by different authors. In the PPG I system, based on Gasper et al. (2016), Blechnum is one of 18 genera in the subfamily Blechnoideae, and has about 30 species. Other sources use a very broadly defined Blechnum s.l., including accepting only two other genera in the subfamily. The genus then has about 250 species. In the PPG I circumscription, the genus is mostly neotropical, with a few southern African species.
Woodwardia fimbriata, known by the common name giant chain fern, is an evergreen perennial fern species in the family Blechnaceae, in the eupolypods II clade of the order Polypodiales, in the class Polypodiopsida. It is native to western North America from British Columbia through California, including the Sierra Nevada, into Baja California.
Struthiopteris spicant, syn. Blechnum spicant, is a species of fern in the family Blechnaceae, known by the common names hard-fern or deer fern. It is native to Europe, western Asia, northern Africa, and western North America. Like some other species in the family Blechnaceae, it has two types of leaves. The sterile leaves have flat, wavy-margined leaflets 5 to 8 millimeters wide, while the fertile leaves have much narrower leaflets, each with two thick rows of sori on the underside.
Salpichlaena is a genus containing two species of fern in the family Blechnaceae. Species of the genus are native to Central and South America. These ferns have climbing leaves with a rachis that twines around tree branches and other supports.
Austroblechnum durum, synonym Blechnum durum, is a species of fern in the family Blechnaceae. The fern is endemic to New Zealand.
Parablechnum wattsii, synonym Blechnum wattsii, is a common terrestrial fern growing in rainforest and open forest. It is often seen near creeks in much of south eastern Australia, including Victoria, Tasmania, South Australia, New South Wales and Queensland. The specific epithet wattsii honours William Walter Watts (1856-1920). Watts was considered an authority on mosses and ferns and has more than 30 species named for him. Common names by which the species may be called are hard water fern - from its stiff leathery fronds, leech fern - as forest workers often encounter leaches while working in clusters of these ferns, hard hill fern - from the fern's habit and habitat, and red cabbage fern - from the bronze-pink colour of the young fronds resembling cooked red cabbage.
Diplazium australe, commonly known as the Austral lady fern, is a small fern occurring in eastern Australia, New Zealand and Norfolk Island. The habitat is moist shaded areas, often occurring in rainforest.
Wessiea is an extinct morphogenus of fern not placed in a specific family. Wessiea is known from Late Cretaceous and Middle Miocene age fossils found in Central Washington USA and Southern Alberta Canada. Two species are described in the morphogenus, W. oroszii and the type species W. yakimaensis.
Osmunda wehrii is an extinct species of fern in the modern genus Osmunda of the family Osmundaceae. Osmunda wehrii is known from Langhian age Miocene fossils found in Central Washington.
Myriopteris tomentosa, formerly known as Cheilanthes tomentosa, is a perennial fern known as woolly lipfern. Woolly lipfern is native to the southern United States, from Virginia to Arizona and Georgia, and Mexico.
Liquidambar changii is an extinct species of sweetgum in the Altingiaceae genus Liquidambar. Liquidambar changii is known from Middle Miocene fossils found in Central Washington.
Quercus hiholensis is an extinct species of oak in the Fagaceae genus Quercus. The species is known from Middle Miocene fossils found in Central Washington.
Shirleya is an extinct genus in the crape myrtle family, Lythraceae, which contains a single species, Shirleya grahamae. The genus and species are known from Middle Miocene fossils found in Central Washington.
Argyrochosma connectens is a small cheilanthoid fern endemic to Sichuan, China. It is the only member of its genus known from Asia. Relatively rare, it is found growing in the crevices of limestone rocks in hot, dry valleys. The species was long classified in the genus Pellaea, but after a phylogenetic study in 2015 was transferred to Argyrochosma.