Campbelloceras

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Campbelloceras
Temporal range: Lower Ordovician
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Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: Nautiloidea
Order: Tarphycerida
Family: Tarphyceratidae
Genus: Campbelloceras
Ulrich and Foerste, 1936

Campbelloceras is a tarphyceratid nautiloid known from the Lower Ordovician, Upper Canadian Epoch of North America, where it is widespread. Campbelloceras was named by Ulrich and Foerste in 1936.

The Tarphyceratidae are tightly coiled, evolute Tarphycerida with ventral siphuncles. The dorsum is characteristically impressed where the whorl presses against the venter of the previous. The Tarphyceratidae are derived from Bassleroceras or possibly from some member of the Estonioceratidae.

Nautiloid subclass of molluscs

Nautiloids are a large and diverse group of marine cephalopods (Mollusca) belonging to the subclass Nautiloidea that began in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Nautiloids flourished during the early Paleozoic era, where they constituted the main predatory animals, and developed an extraordinary diversity of shell shapes and forms. Some 2,500 species of fossil nautiloids are known, but only a handful of species survive to the present day.

The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.2 million years from the end of the Cambrian Period 485.4 million years ago (Mya) to the start of the Silurian Period 443.8 Mya.

The shell of Campbelloceras has a circular whorl section, only slightly impressed, and a siphuncle that is close to the venter in all growth stages. Campbelloceras differs from Tarphyceras in that the rate of expansion is greater, the siphuncle is proportionally largers, and an impression is shallower.

Siphuncle

The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and Spirula. In the case of the cuttlefish, the siphuncle is indistinct and connects all the small chambers of that animal's highly modified shell; in the other cephalopods it is thread-like and passes through small openings in the walls dividing the chambers.

Tarphyeras is a genus of tarphyceratid with whorls rounded in cross section, having a deeply impressed dorsum and a ventral to subcentral siphuncle, known from the Lower Ord of North America. It differs from Campbelloceras in that Campbelloceras is only slightly impressed, from Centrotarphyceras in that Centrotarphyceras is subquadrate and has a central siphuncle, and from Trocholites in that although Trocholites is subcircular in cross section, the siphuncle is subdorsal.

Campbelloceras may have given rise to the barrandeocerid Plectoceras (Plectoceratidae) through simplification of the connecting rings, and to Tarphyceras through tighter coiling, development of a deeper impression, and reduction in the size of the siphuncle. Campbelloceras may be conceivably derived from Estonioceras (Estonioceratidae). The cross section of Campbelloceras and that of the outer whorl of Estonioceras are similar, except for Campbelloceras being slightly impressed. Both have proportionally large siphuncles, near the venter.

Plectoceras is a genus of nautiloids included in the tarphycerid suborder Barrandeocerina that lived during the Middle and Late Ordovician. It has been found widespread in the Middle and Upper Ordovician of North America.

The Plectoceratidae is a family of tarphycerids in the suborder Barrandeocerina established as a place for the genus Plectoceras; defined simply as coiled, costate barrandeocerids with subcentral adult siphuncle.

<i>Estonioceras</i> genus of molluscs

Estonioceras is an extinct genus of tarphyceridan nautiloids from the Ordovician of Europe.

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<i>Aphetoceras</i> genus of molluscs

Aphetoceras is a genus of Tarphycerida cephalopod within the Estonioceratidae; loosely coiled without an impression along the dorsal margin; early whorls barely teaching, separating then diverging in the final mature whorl; weakly ribbed in some. The cross section of Aphetoceras is higher than wide, making it compressed in form. The dorsum, along the inner curve, is more broadly rounded than the venter which lies along the outer curve. The siphuncle is relatively large, located near but not at the ventral margin; lined with secondary deposits. Chambers are empty.

Trocholites is a tarphycerid genus in the family Trocholitidae from the Middle and Upper Ordovician with a gradually expanding, weakly ribbed shell; whorls in contact, dorsum slightly impressed; cross section depressed, venter and sides rounded; siphuncle close to but not at the dorsal margin.

<i>Discoceras</i> genus of molluscs

Discoceras is an extinct marine cephalopod mollusk, a member of the Trocholitidae in the Tarphycerida. It is distinct from Discosorus, It is characterized by closely coiled, gradually expanding shells with a subquadrate cross section, that may be ribbed or smooth. The sides are broadly rounded; the venter is wide and slightly rounded. The maximum width is slightly dorsal of the middle. The dorsum has a slight to moderate impression. The siphuncle starts off central for the first half whorl then becomes marginodorsal in the succeeding two whorls, then subdorsal at maturity. As with Trocholites, the dorsal siphuncle in Discoceras probably indicates an orientation during life that places the back of the living chamber high in the shell.

<i>Clymenia</i> (ammonite) genus of molluscs (fossil)

Clymenia is a genus in the ammonoid order Clymeniida, restricted to the Upper Devonian, characterized as with all clymeniids by a dorsal siphuncle that runs along the inside of the whorls, unusual for ammonoids.

The Tarphycerida were the first of the coiled cephalopods, found in marine sediments from the Lower Ordovician to the Middle Devonian. Some, such as Aphetoceras and Estonioceras, are loosely coiled and gyroconic; others, such as Campbelloceras, Tarphyceras, and Trocholites, are tightly coiled, but evolute with all whorls showing. The body chamber of tarphycerids is typically long and tubular, as much as half the length of the containing whorl in most, greater than in the Silurian Ophidioceratidae. The Tarphycerida evolved from the elongated, compressed, exogastric Bassleroceratidae, probably Bassleroceras, around the end of the Gasconadian through forms like Aphetoceras. Close coiling developed rather quickly, and both gyroconic and evolute forms are found in the early middle Canadian.

Curtoceras is a genus in the tarphycerid family Trocholitidae found widespread in the late Early and Middle Ordovician of North America and northern Europe. Curtoceras has a shell that is gradually expanded, with half the fully mature body chamber divergent from the preceding volution. Whorl sections are near equidimentional with the inner margin (dosum) moderately impressed. The surface may be smooth or weakly ribbed. The siphuncle is ventral in the initial chamber and becomes dorsal after one volution. With the exception of the dorsal siphuncle, Curtoceras is somewhat similar to the tarphyceratid Campbelloceras

The Trocholitidae are Tarphycerida with whorls in close contact as with the Tarphyceratidae, but in which the siphuncle, similar in structure, becomes dorsal. The Trocholitidae are derived from the Tarphyceratidae, perhaps from different tarphyceratids.

Trigonocerataceae family of molluscs

The Trigonocerataceae are a superfamily within the Nautilida that ranged from the Devonian to the Triassic, thought to have contained the source for the Nautilaceae in which Nautilus is found.

Syringonautilidae is a family of Nautiloidea from the middle to late Triassic. Syringonautilidae comprise the last of the Trigonocerataceae and are the source for the Nautilaceae which continued the Nautiloidea through the Mesozoic and into the Cenozoic right down to the recent. Syringonautilidae is a strictly Triassic family, derived early in the Triassic from the Grypoceratidae.

Grypoceratidae family of molluscs

Grypoceratidae is the longest-lived family of the Trigonocerataceae, or of the near equivalent Centroceratina; members of the Nautilida from the Upper Paleozoic and Triassic.

The Centroceratidae is the ancestral family of the Trigonocerataceae and of the equivalent Centroceratina; extinct shelled cephalopods belonging to the order Nautilida

Aphelaeceras is an extinct genus from the nautilid family Trigonoceratidae which is part of the Trigonocerataceae, that lived during the Mississippian Period in the late Paleozoic.

Thuringionautilus is a genus of large, moderately involute, nautiloids from the nautilid family Tainoceratidae. The whorl section is subquadrate, flanks slightly convex, venter broad with a median furrow. Ventral shoulders, narrowly rounded to subangular; umbilical shoulders, broadly rounded. Longitudinal nodes slope diagonally backwards on the venter toward the furrow. Suture, slightly sinuous. Siphuncle, subdorsal.

The Tetragonoceratidae is a small family of nautilitids constituting a part of the superfamily Tainocerataceae in which shells are coiled with a generally quadrate whorl section. Coiling is either gyroconic or evoluute with a slight dorsal impression. Flanks diverge from the umbilical to the ventral shoulders so as to make the whorl sections widest close to the venter. Nodes made develop on the flanks and shoulders. The siphuncle is tubular, typically found near the venter. Sutures are with lateral lobes and may possess dorsal and ventral lobes as well.

Chidleyenoceras is a Middle Ordovician tarphyceroid with a closely coiled, evolute shell; whorl section subquadrate, widest just above a broadly rounded venter; dorsum with a broad shallow impression; sutures moderately spaced, weakly sinuous; siphuncle large, subventral, apparently orthochoantitic with tubular segments.

<i>Domatoceras</i> genus of molluscs

Domatoceras is a nautiloid genus and member of the Grypoceratidae from the Pennsylvanian and Permian with a wide spread distribution.

Aethoceras is a genus of Tarphycerida nautiloids included in the family Estonioceratidae for which the shell is a loosely coiled, gradually expanding dextral torticone with a slightly derpressed whorl section. Siphuncle small, ventral, submarginal. Whorl section somewhat resembles early stages of Estonioceras in being laterally flanged. Trochoidal coiling brings to mind the later Trocholitidae. Aethoceras was originally found in the lower Ordovician of Western Australia.

References

    The Treatise on Invertebrate Paleontology published by the Geological Society of America and the University of Kansas Press, is a definitive multi-authored work of some 50 volumes, written by more than 300 paleontologists, and covering every phylum, class, order, family, and genus of fossil and extant invertebrate animals. The prehistoric invertebrates are described as to their taxonomy, morphology, paleoecology, stratigraphic and paleogeographic range. However, genera with no fossil record whatsoever have just a very brief listing.