Carcharodon hubbelli | |
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Holotype jaws and teeth (UF 226255), also known as the Sacaco specimen | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Chondrichthyes |
Subclass: | Elasmobranchii |
Subdivision: | Selachimorpha |
Order: | Lamniformes |
Family: | Lamnidae |
Genus: | Carcharodon |
Species: | †C. hubbelli |
Binomial name | |
†Carcharodon hubbelli Ehret et al., 2012 | |
Carcharodon hubbelli, also known as Hubbell's white shark, is an extinct species of white shark that evolved between 8 and 5 million years ago during the Late Miocene to Early Pliocene epochs. This shark is a transitional species, showing intermediate features between the extant great white shark and the fossil white shark, C. hastalis . [2] [3] [4] C. hubbelli appears to be geographically restricted to the Pacific Ocean, with fossils of C. hubbelli recovered from Peru, [4] Chile [ citation needed ], California, [5] and New Zealand. [6] This exclusive distribution suggests a Pacific origin for the great white shark.
This shark was named in honor of Dr. Gordon Hubbell (the scientist who recovered the specimen from a farmer who found it in 1988) [7] in recognition of his contribution to shark paleontology and for donating the specimen to the Florida Museum of Natural History in 2009. [8] [3] [6] It was about the size of the modern great white shark, reaching 4.9–5.1 metres (16–17 ft) long. [3] [9] Its growth curve shows that it grew at a slower rate than the modern great white shark. [3]
Only one associated dentition of C. hubbelli is known (UF 226255), with a total of 222 teeth (and 45 vertebral centra). It was recovered from the Pisco Formation of Southern Peru. The tooth morphology is most similar to the extant great white, in having labiolingually flattened triangular cusps with serrated edges. The dentition differs from the extant species in having a distally-inclined upper intermediate tooth (it is mesially-inclined in the great white). The serrations are also qualitatively different, being finer and diminish in size apically (towards the tip). [10]
As a transitional species, C. hubbelli possesses a mixture of features present in extinct white sharks like C. hastalis and in the extant white shark, C. carcharias. The distal inclination of the intermediate tooth is characteristic of C. hastalis, but the presence of serrations (though not fully developed) and a second upper anterior tooth larger than the lower second anterior tooth are characteristic of C. carcharias. [10]
The development of serrated edges on the teeth of C. hubbelli suggest a shift in its diet to rely more on marine mammals as in C. carcharias. The bone of a scavenged mysticete whale with a tooth of C. hubbelli embedded within it supports this hypothesis. [11]
The great white shark, also known as the white shark, white pointer, or simply great white, is a species of large mackerel shark which can be found in the coastal surface waters of all the major oceans. It is the only known surviving species of its genus Carcharodon. The great white shark is notable for its size, with the largest preserved female specimen measuring 5.83 m (19.1 ft) in length and around 2,000 kg (4,400 lb) in weight at maturity. However, most are smaller; males measure 3.4 to 4.0 m, and females measure 4.6 to 4.9 m on average. According to a 2014 study, the lifespan of great white sharks is estimated to be as long as 70 years or more, well above previous estimates, making it one of the longest lived cartilaginous fishes currently known. According to the same study, male great white sharks take 26 years to reach sexual maturity, while the females take 33 years to be ready to produce offspring. Great white sharks can swim at speeds of 25 km/h (16 mph) for short bursts and to depths of 1,200 m (3,900 ft).
The Lamniformes are an order of sharks commonly known as mackerel sharks. It includes some of the most familiar species of sharks, such as the great white, as well as more unusual representatives, such as the goblin shark and megamouth shark.
Carcharodon is a genus of sharks within the family Lamnidae, colloquially called the "white sharks." The only extant member is the great white shark. The extant species was preceded by a number of fossil (extinct) species including C. hubbelli and C. hastalis. The first appearance of the genus may have been as early as the Early Miocene or Late Oligocene. Carcharocles megalodon is still argued by some paleontologists to be a close relative of Carcharodon carcharias - as well as being in the same genus. The megalodon's scientific name was originally "Carcharodon" megalodon, but more recently, the giant shark has been assigned by most scientists to either the genus Carcharocles or Otodus.
Otodus megalodon, commonly known as megalodon, is an extinct species of giant mackerel shark that lived approximately 23 to 3.6 million years ago (Mya), from the Early Miocene to the Pliocene epochs. O. megalodon was formerly thought to be a member of the family Lamnidae and a close relative of the great white shark, but has been reclassified into the extinct family Otodontidae, which diverged from the great white shark during the Early Cretaceous.
The Lamnidae are the family of mackerel sharks known as white sharks. They are large, fast-swimming predatory fish found in oceans worldwide, though prefer environments with colder water. The name of the family is formed from the Greek word lamna, which means "fish of prey", and was derived from the Greek legendary creature, the Lamia.
Otodus is an extinct, cosmopolitan genus of mackerel shark which lived from the Paleocene to the Pliocene epoch. The name Otodus comes from Ancient Greek ὠτ- and ὀδούς – thus, "ear-shaped tooth".
Isurus is a genus of mackerel sharks in the family Lamnidae, commonly known as the mako sharks. They are largely pelagic, fast predatory fish capable of swimming at speeds up to 50 km/h (31 mph).
Otodus angustidens is an extinct species of prehistoric megatoothed sharks in the genus Otodus, which lived during the Oligocene and Miocene epochs about 33 to 22 million years ago. The largest individuals were about 11–12 metres (36–39 ft) long. This shark is related to another extinct megatoothed shark, the famous Otodus megalodon.
Thalassocnus is an extinct genus of semiaquatic ground sloths from the Miocene and Pliocene of the Pacific South American coast. It is monotypic within the subfamily Thalassocninae. The five species—T. antiquus, T. natans, T. littoralis, T. carolomartini, and T. yuacensis—represent a chronospecies, a population gradually adapting to marine life in one direct lineage. They are the only known aquatic sloths, but they may have also been adapted to a terrestrial lifestyle. They have been found in the Pisco Formation of Peru, the Tafna Formation of Argentina, and the Bahía Inglesa, Coquimbo, and Horcón formations of Chile. Thalassocninae has been placed in both the families Megatheriidae and Nothrotheriidae.
The Mayoan age is a period of geologic time from 11.8 to 10 Ma, within the Middle to Late Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Laventan and precedes the Chasicoan age.
Piscogavialis is an extinct monospecific genus of gryposuchine gavialid crocodylian. The only species yet known is P. jugaliperforatus. Fossils of Piscogavialis have been found from the Mio-Pliocene Pisco Formation of the Sacaco Basin in southern Peru in 1998, where it coexisted with the much smaller gavialid Sacacosuchus.
Otodus chubutensis, meaning "ear-shaped tooth of Chubut", from Ancient Greek ὠτ and ὀδούς – thus, "ear-shaped tooth", is an extinct species of prehistoric megatoothed sharks in the genus Otodus, that lived during Early Miocene to Middle Miocene. The largest individuals were about 13.5 metres (44 ft) long. This shark is considered a close relative of the famous prehistoric megatoothed shark O. megalodon. However, as is the case with O. megalodon, the classification of this species is disputed.
Otodontidae is an extinct family of sharks belonging to the order Lamniformes. Its members have been described as megatoothed sharks. They lived from the Early Cretaceous to the Pliocene, and included genera such as Otodus, including the giant megalodon. Recent studies of the newly described genus Megalolamna indicate that the members of the genus Carcharocles should be reclassified as members of the genus Otodus. The genus Cretalamna which lived from the mid-Cretaceous-Paleogene is believed to be directly ancestral to Otodus, and thus to megalodon.
Livyatan is an extinct genus of macroraptorial sperm whale containing one known species: L. melvillei. The genus name was inspired by the biblical sea monster Leviathan, and the species name by Herman Melville, the author of the famous novel Moby-Dick about a white bull sperm whale. Herman Melville often referred to whales as "Leviathans" in his book. It is mainly known from the Pisco Formation of Peru during the Tortonian stage of the Miocene epoch, about 9.9–8.9 million years ago (mya); however, finds of isolated teeth from other locations such as Chile, Argentina, the United States (California), South Africa and Australia imply that either it or a close relative survived into the Pliocene, around 5 mya, and may have had a global presence. It was a member of a group of macroraptorial sperm whales and was probably an apex predator, preying on whales, seals and so forth. Characteristically of raptorial sperm whales, Livyatan had functional, enamel-coated teeth on the upper and lower jaws, as well as several features suitable for hunting large prey.
Cretalamna is a genus of extinct otodontid shark that lived from the latest Early Cretaceous to Eocene epoch. It is considered by many to be the ancestor of the largest sharks to have ever lived, such as Otodus angustidens, Otodus chubutensis, and Otodus megalodon.
The Temblor Formation is a geologic formation in California. It preserves fossils dating back from the Late Oligocene to the Middle Miocene of the Neogene period. It is notable for the famous Sharktooth Hill deposit.
The Pisco Formation is a geologic formation located in Peru, on the southern coastal desert of Ica and Arequipa. The approximately 640 metres (2,100 ft) thick formation was deposited in the Pisco Basin, spanning an age from the Late Miocene up to the Early Pliocene, roughly from 9.6 to 4.5 Ma. The tuffaceous sandstones, diatomaceous siltstones, conglomerates and dolomites were deposited in a lagoonal to near-shore environment, in bays similar to other Pacific South American formations as the Bahía Inglesa and Coquimbo Formations of Chile.
Cosmopolitodus is an extinct genus of mackerel shark that lived between thirty and one million years ago during the late Oligocene to the Early Pleistocene epochs. Its type species is Cosmopolitodus hastalis, the broad-tooth mako. In 2021, Isurus planus was reassigned to the genus, and thus became the second species C. planus. However, some researchers still consider both species of Cosmopolitodus as species of Carcharodon.
Carcharomodus escheri, commonly nicknamed the serrated mako shark or Escher's mako shark, is an extinct lamnid that lived during the Miocene. It has been formerly thought to have been the transitional between the broad-toothed "mako" Cosmopolitodus hastalis and the modern great white, but is now considered to be an evolutionary dead-end with the discovery of Carcharodon hubbelli. Fossil examples have been found along northern Atlantic coastlines and in parts of Western and Central Europe.
Alopias palatasi, commonly referred to as the serrated giant thresher, is an extinct species of giant thresher shark that lived approximately 20.44 to 13.7 million years ago during the Miocene epoch, and is known for its uniquely serrated teeth. It is only known from such isolated teeth, which are large and can measure up to an excess of 4 centimetres (2 in), equating to a size rivaling the great white shark, but are rare and found in deposits in the East Coast of the United States and Malta. Teeth of A. palatasi are strikingly similar to those of the giant thresher Alopias grandis, and the former has been considered as a variant of the latter in the past. Scientists hypothesized that A. palatasi may have had attained lengths comparable with the great white shark and a body outline similar to it.