Cheirothricidae Temporal range: | |
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Fossil of Cheirothrix lewisii | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Order: | Aulopiformes (?) |
Family: | † Cheirothricidae Woodward, 1901 |
Genera | |
See text |
Cheirothricidae [1] [2] [3] is a family of extinct marine ray-finned fish, perhaps belonging to the Aulopiformes, although they are tentatively placed as indeterminate eurypterygians. [4] They lived during the Upper Cretaceous (Cenomanian - Campanian, about 95 - 71 million years ago) and their fossil remains are found in the Middle East and Europe. They were characterized by very expanded even fins.
The main characteristic of these small fish (the length of the body was usually between 3 and 20 centimeters in length) was given by the extreme development of the even fins; in Cheirothrix species [5] the pectoral fins were particularly developed, while in Telepholis and Exocoetoides also the pelvic fins were large. In any case, the fins were large and provided with very long rays.
Cheirothrix was the largest genus, had a more robust body and could reach 20 centimeters in length; Telepholis, usually shorter and with a thinner body, had shorter pectoral fins than Exocoetoides and had some small and thin shields in the dorsal region, rounded or polygonal, adorned in the center with a tubercle. The caudal fin was not very indented. However, Telepholis may not be a cheirothricid. [4]
Exocoetoides, on the other hand, was decidedly small in size (between 3 and 7 centimeters in length) and the pectoral fins were particularly long (they could reach the anal region); in the caudal fin both lobes were the same size.
Established in 1901 by Arthur Smith Woodward, the family Cheirothricidae was widespread in the ancient Tethys Ocean, in the areas currently occupied by the Middle East and Europe.
Two genera are known in the Cretaceous of Lebanon (Cheirothrix in deposits of the Santonian, and Exocoetoides from the Cenomanian), and in Germany (Cheirothrix and Telepholis from the Campanian). [6] [7]
Members of this family are generally considered to be representatives of the Aulopiformes [8] ; in particular, affinities have been proposed with the extinct group of Enchodontidae. However, a 2009 study found them to be indeterminate members of the Eurypterygii. [4]
The Cheirothricidae had a morphology remarkably similar to that of the extant flying fishes and to that of other extinct forms like Thoracopterus . Like today's Cypselurus and Exocoetus , Cheirothricidae were also likely to perform a sort of gliding flight over water, thanks to the notable expansion of the pectoral and pelvic fins.[ citation needed ]
The Tetraodontiformes are an order of highly derived ray-finned fish, also called the Plectognathi. Sometimes these are classified as a suborder of the order Perciformes. The Tetraodontiformes are represented by 10 extant families and at least 349 species overall; most are marine and dwell in and around tropical coral reefs, but a few species are found in freshwater streams and estuaries. They have no close relatives, and descend from a line of coral-dwelling species that emerged around 80 million years ago.
Aulopiformes is a diverse order of marine ray-finned fish consisting of some 15 extant and several prehistoric families with about 45 genera and over 230 species. The common names grinners, lizardfishes and allies, or aulopiforms are sometimes used for this group. The scientific name means "Aulopus-shaped", from Aulopus + the standard fish order suffix "-formes". It ultimately derives from Ancient Greek aulós + Latin forma, the former in reference to the elongated shape of many aulopiforms.
The Congridae are the family of conger and garden eels. Congers are valuable and often large food fishes, while garden eels live in colonies, all protruding from the sea floor after the manner of plants in a garden. The family includes over 180 species in 32 genera.
Enchodus is an extinct genus of aulopiform ray-finned fish related to lancetfish and lizardfish. Species of Enchodus flourished during the Late Cretaceous, and there is some evidence that they may have survived to the Paleocene or Eocene; however, this may just represent reworked Cretaceous material.
The cusk-eel family, Ophidiidae, is a group of marine bony fishes in the Ophidiiformes order. The scientific name is from the Greek ophis meaning "snake", and refers to their eel-like appearance. True eels diverged from other ray-finned fish during the Jurassic, while cusk-eels are part of the Percomorpha clade, along with tuna, perch, seahorses and others.
Berycopsis is an extinct genus of marine ray-finned fish from the Late Cretaceous period. Fossils are known from England, Germany, and Lebanon. A potential specimen is known from the Czech Republic.
Protosphyraena is a fossil genus of swordfish-like marine fish, that thrived worldwide during the Upper Cretaceous Period (Coniacian-Maastrichtian). Though fossil remains of this taxon have been found in both Europe and Asia, it is perhaps best known from the Smoky Hill Member of the Niobrara Chalk Formation of Kansas. Protosphyraena was a large fish, averaging 2–3 metres in length. Protosphyraena shared the Cretaceous oceans with aquatic reptiles, such as mosasaurs and plesiosaurs, as well as with many other species of extinct predatory fish. The name Protosphyraena is a combination of the Greek word protos ("early") plus Sphyraena, the genus name for barracuda, as paleontologists initially mistook Protosphyraena for an ancestral barracuda. Recent research shows that the genus Protosphyraena is not at all related to the true swordfish-family Xiphiidae, but belongs to the extinct family Pachycormidae.
Apateodus is a genus of prehistoric marine ray-finned fish which was described by Woodward in 1901. It was a relative of modern lizardfish and lancetfish in the order Aulopiformes, and one of a number of prominent nektonic aulopiforms of Cretaceous marine ecosystems.
Apateopholis is an extinct genus of prehistoric marine ray-finned fish. It is the only member of the family Apateopholidae and contains a single species, A. laniatus, from the Cenomanian of Lebanon. It was a relative of modern lizardfish and lancetfish in the order Aulopiformes. At least one study has found it to be a potential sister genus to the enigmatic Yabrudichthys of the West Bank.
Cheirothrix is an extinct genus of marine ray-finned fish from the Late Cretaceous. It is named for its enlarged pectoral fins, which show close convergence with modern flying fish.
Exocoetoides is an extinct genus of prehistoric ray-finned fish.
Carsothrissops is an extinct genus of prehistoric marine ray-finned fish that lived during the Cenomanian. It contains a single species, C. delorenzi from Komen, Slovenia.
Barcarenichthys is an extinct genus of prehistoric marine ray-finned fish that lived during the Late Cretaceous. It contains a single species, B. joneti, from the late Cenomanian of Portugal.
Ampheristus is an extinct genus of prehistoric marine ray-finned fish. It was a basal or stem member of the family Ophidiidae, which contains modern cusk-eels. Fossils are known from worldwide from the Late Cretaceous to the late Paleogene, making it a rather successful survivor of the Cretaceous-Paleogene extinction event.
Anomoeodus is an extinct genus of prehistoric marine ray-finned fish belonging to the family Pycnodontidae. This genus lived during the Late Cretaceous period, ranging from the Albian to Maastrichtian ages, and had a wide geographic distribution, with fossils found in France, Belgium, the Netherlands, Spain, Egypt, Uzbekistan, and the United States. Some studies have recovered it as a wastebasket taxon. The first fossils of Anomoeodus were described by Louis Agassiz in 1833, although they were described under Pycnodus.
Boreosomus is an extinct genus of Triassic marine ray-finned fish. It was first described from the Arctic island of Spitsbergen, hence its genus name, but was later also discovered in other parts of the world. The type species is Boreosomus arcticus.
Charitosomus is an extinct genus of prehistoric marine ray-finned fish from the Late Cretaceous, related to modern beaked salmons. They were nektonic carnivores in life.
Sorbinichthys is an extinct genus of clupeomorph bony fish from the Cenomanian of Lebanon and Morocco.
Tselfatiiformes is an extinct order of bony fishes from the infraclass Teleostei. The order represents the most important radiation of marine teleosts during the Cretaceous period. Fossils of tselfatiiforms are known from Europe, North America, central and northern South America, the Middle East and North Africa.
Halecidae is an extinct family of aulopiform fish known from the Cretaceous. It is one of the Enchodontoidei, a diverse group of aulopiforms that were dominant marine fish during the Cretaceous.
Note: This article has been expanded using material based on a translation of an article from the Italian Wikipedia.