| Green liveforever | |
|---|---|
 | |
| Dudleya virens ssp. hassei | |
Scientific classification  | |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Clade: | Angiosperms |
| Clade: | Eudicots |
| Order: | Saxifragales |
| Family: | Crassulaceae |
| Genus: | Dudleya |
| Species: | D. virens |
| Binomial name | |
| Dudleya virens | |
| Synonyms [1] | |
| |
Dudleya virens, the green liveforever or bright green dudleya, is an uncommon species of perennial, succulent plant in the family Crassulaceae, native to several coastal southern California and Baja California locations. [2]
The succulent is found growing in the Palos Verdes Hills on the Palos Verdes Peninsula coast in Los Angeles County; on several of the Channel Islands in California; and on Guadalupe Island off the northwestern Baja California Peninsula, Mexico.
Dudleya virens leaves are fleshy and strap-shaped, 8–20 cm long and 1.5–3 cm broad, tapering from the base (or from near middle) and are mostly green. They are arranged in a rosette.
The flowers are white, with five petals 7–10 mm long. They are produced in April, May, and June. Hummingbirds visit the flowers for their nectar.
| Name | Authority | Common name | Distribution | Image |
|---|---|---|---|---|
| Species Dudleya virens (Rose) Moran | ||||
| D. v. subsp. extima | ||||
| subsp. extima | Moran | Guadalupe green liveforever | Endemic to Guadalupe Island | |
| Caudex 1 to 2.5 cm thick, either branching into clumps of up to 15 rosettes or elongating up to 30 cm. Rosettes are 5 to 15 cm wide, containing 15 to 55 crowded leaves. Leaves are bright green or glaucous. Leaf shape is linear to oblong-oblanceolate, 4 to 7 or rarely up to 17 cm long, 6 to 10 mm wide above when linear but up to 15 mm wide above when oblanceolate, 8 to 15 mm wide at the base and 2 to 4 mm thick. Peduncle 5 to 20 cm tall, 2 to 5 mm thick, inflorescence 2 to 7 cm wide, composed of 2 to 4 ascending branches. Flowers are white with petals 7 to 10 mm long. Chromosomes n = 17, 34. [3] [4] | ||||
| D. v. subsp. hassei | ||||
| subsp. hassei syn. Stylophyllum hassei syn. Cotyledon hassei syn. Echeveria hassei syn. Dudleya hassei | (Rose) Moran [3] Rose in Britton & Rose [5] (Rose) Fedde [6] (Rose) Berger [7] (Rose) Moran [8] | Catalina Island liveforever | Endemic to Santa Catalina Island. |  |
| Caudex 1 to 3 cm thick, branching into clumps 1 m wide. Rosettes are 4 to 8 cm wide, containing 15 to 30 erect leaves. Leaves are generally glaucous, but rarely can be green. Leaf shape is linear-oblong to linear-lanceolate and 5 to 10 cm long, 2 to 4 mm thick. Peduncle 10 to 30 cm tall, inflorescence branching into 2 to 4 close set or once bifurcate branches. Flowers are white with petals 8 to 10 mm long. Chromosomes n = 34. [9] [4] | ||||
| D. v. subsp. insularis | ||||
| subsp. insularis syn. Stylophyllum insulare syn. Cotyledon insularis syn. Cotyledon viscida var. insularis syn. Echeveria insularis syn. Echeveria viscida var. insularis syn. Dudleya insularis | (Rose) Moran Rose in Britton & Rose [5] (Rose) Fedde [6] (Rose) S. Watson [10] (Rose) Berger [7] (S. Watson) Berger [11] (Rose) P. H. Thomson [12] | Island green dudleya | Occurs in the Palos Verdes Hills, Santa Catalina and San Nicolas islands. |  |
| Caudex 2 to 6 cm thick, up to 1 m long, branching into clumps 2 m wide. Rosettes are 10 to 25 cm wide, containing 20 to 50 spreading to erect leaves. Leaves sometimes glaucous, sometimes green. Leaf shape is triangular-lanceolate, 6 to 25 cm long and 10 to 32 mm wide above, 2 to 4 cm wide at the base, with an obtuse tip. Peduncle 6 to 70 cm tall, 5 to 15 mm thick. Flowers are white with red to orange keels, petals 8 to 10 mm long. Chromosomes n = 17. [13] [4] | ||||
| D. v.subsp. virens | ||||
| subsp. virens syn. Stylophyllum virens syn. Cotyledon virens syn. Echeveria virens syn. Dudleya virens syn. Stylophyllum albida syn. Cotyledon albida syn. Echeveria albida syn. Dudleya albida | (Rose) Moran Rose in Britton & Rose [5] (Rose) Fedde [6] (Rose) Berger [7] (Rose) Moran [14] Rose in Britton & Rose [5] (Rose) Fedde [6] (Rose) Berger [7] (Rose) P. H. Thomson [12] | Bright green dudleya Alabaster plant | Endemic to San Clemente Island. |  |
| Caudex 1 to 3.2 cm wide, branching into clumps at least 40 cm wide. Rosettes are 5 to 10 cm wide, containing 20 to 50 leaves. Leaves are usually green, rarely glaucous. Leaf shape is triangular-lanceolate to linear-lanceolate, 3 to 10 cm long and 10 to 15 mm wide above, 1.5 to 3 cm wide at the base, with an acute tip. Peduncle is 7 to 46 cm long, 4 to 7 mm thick, inflorescence 6 to 15 cm wide with 3 to 4 close-set branches towards the apex. Flowers are white with red to orange keels, petals 8 to 10 mm long. Chromosomes n = 17. [15] [4] | ||||
There has been some developing arguments on how to best categorize the various subspecies of Dudleya virens. The division stems from the fact that the current groupings of the Dudleya virens subspecies are polyphyletic as opposed to monophyletic [16] . The overall evolutionary history of Dudleya virens has been difficult to pin down [17] which has contributed to the confusion surrounding the clarification of the subspecies. This has lead to some researchers suggesting that the white leafed sub species of Dudleya virens should be separated into its own species that is distinct from the green colored subspecies of Dudleya virens, although this theory is not backed with any DNA results [16] .
Dudleya virens is cultivated as an ornamental plant in the specialty native plants and succulents horticulture trade. [18] It is used in containers, drought tolerant landscapes, and habitat gardens. It prefers well-draining soil, and grows best in full sun or light shade.
Dudleya virens has been specifically affected by the surge in Dudleya poaching to be sold in a similar way to Dudleya farinosa which was covered in the 2021 documentary "Plant Heist" [19] . Where Dudleya virens plants are illegally harvested and shipped back to regions of Asia to be sold for profit as medicine or a plant [19] . The succulent is also threatened by invasive grazing animal species that were introduced during the colonization of the Americas [4] . These animals over consume the plants [4] . The poaching [19] by people and predation [4] by invasive species has led to Dudleya virens, specifically Dudleya virens spp. insularis, to be ranked a 1B.2 in terms of rarity (rare, threatened, or endangered) by the state of California [20] .
Through consorted conservation efforts, Dudleya virens has been able to make a considerable ecological come back from its previously concerning status [4] . One of the major factors in their successful rehabilitation has been the removal of invasive grazing species: rabbits, horse, and pigs that were introduced during the colonization of California. These animals tend to over consume several Dudleya virens subspecies throughout the Channel Islands [4] . The other major factor is that California passed a bill that makes poaching Dudleya virens with an intent to sell a fineable offense [19] [20] . The removal of these invasive grazers and the crack down of succulent poaching [19] [4] [20] has created the conditions necessary for Dudleya virens to recover across the Channel Islands [4] .
Dudleya, commonly known as liveforevers is a genus of rosette-forming succulent plants in the stonecrop family, Crassulaceae, consisting of about 68 taxa in southwestern North America and Guadalupe Island. The species come in many forms, some large and evergreen, others geophytic and deciduous. Yet, despite their dramatic variations in appearance, most species readily hybridize. The flowers of Dudleya have parts numbered in fives, with the petals arranged in tubular, star-shaped, and bell-shaped forms and, when fruiting, are filled with tiny, ovoid-crescent-shaped seeds.
Dudleya abramsiisubsp. setchellii, known by common name as the Santa Clara Valley dudleya or Santa Clara Valley liveforever, is a member of the Dudleya genus of succulent perennials, members of the family Crassulaceae. The Santa Clara Valley dudleya, endemic to the Santa Clara Valley region in the southern San Francisco Bay Area, was listed on 3 February 1995, as an endangered species. It is considered to be a subspecies of Dudleya abramsii, but its taxonomic status is still unclear. Its closest relative is Dudleya cymosa subsp. paniculata, which is a morphologically similar sister taxon.
Dudleya cymosa is a species complex of evergreen and deciduous succulent plants in the family Crassulaceae known by the common name: canyon liveforever. It is a loosely defined polyphyletic species with a diverse number of subspecies, varying highly in morphology, distribution, and habitat.
Dudleya blochmaniae is a summer-deciduous succulent plant known by the common names Blochman's liveforever or Blochman's dudleya. This species of Dudleya survives part of the year with no aboveground presence, surviving as underground corm-like roots in deciduous months. It is characterized by white, star-shaped and spreading flowers that emerge after sufficient rainfall. It is found along the Pacific coast of the California Floristic Province, from the vicinity of San Luis Obispo in California to Punta Colonet in Baja California.
Dudleya abramsiisubsp.calcicola is a succulent plant known by the common name limestone liveforever, or limestone dudleya. It is endemic to California, where it is a rare resident of limestone outcroppings and rocky slopes in the southern Sierra Nevada and nearby mountains and foothills. It was formerly regarded as Dudleya calcicola.
Dudleya candelabrum is a species of succulent plant known by the common names candleholder liveforever or candleholder dudleya. Endemic to California, this species grows wild only on the northern Channel Islands, where it is found in open rocky places and north-facing slopes. It is characterized by thin, spade-shaped green leaves and an inflorescence covered in long, reflexed bracts, with pale yellow flowers. It has been threatened by poachers shipping plants to South Korea.
Dudleya pulverulenta is a species of perennial succulent plant known by the common names chalk lettuce, chalk dudleya, and chalk liveforever. It is one of the largest Dudleya, with a silvery, waxy rosette that may greatly contrast with its habitat. It is also regarded as one of the most distinctive members of the Dudleya, with one of the most specialized inflorescences in the genus, adapted to hummingbird pollination through its red pendent flowers, the longest corolla, and the highest nectar output. Dudleya pulverulenta has the largest range of all Dudleya, over 1,000 kilometres (620 mi), being found from southern Monterey County in California to the Sierra de San Borja in southern Baja California. It is closely related to Dudleya arizonica, a smaller desert species that tends to lack the specialized floral traits, and Dudleya anthonyi, which differs in a few morphological traits and is restricted to the San Quintín Volcanic Field.
Dudleya traskiae is a rare succulent plant known by the common name Santa Barbara Island liveforever. This Dudleya is endemic to Santa Barbara Island, one of the Channel Islands of California, where it grows on rocky bluffs. The plant has a basal rosette of flat, spade-shaped fleshy leaves up to 15 centimeters long, which are pale green to yellowish. It erects tall stems bearing dense, rounded inflorescences of many bright yellow flowers.
Dudleya greenei is a perennial species of succulent plant known by the common names Greene's liveforever, or Greene's dudleya. It is endemic to the Channel Islands of California, where it grows along the cliffs of four of the eight islands. It is a highly variable plant, presenting with multiple forms and varying levels of ploidy. Taxonomically, this species is an insular segregate of Dudleya caespitosa, and was placed as a stopgap taxon by Reid Moran in his 1951 thesis on the genus. It is characterized by white or green leaf rosettes, loomed over by inflorescences bearing pale yellow to white flowers. It is a member of the subgenus Dudleya, as it cannot be propagated from leaf cuttings, does not grow from a corm, and has tight petals.
Dudleya abramsii is a species complex of succulent plants native to California and parts of Baja California. There are numerous subspecies, some critically endangered, with varying habits and lifestyles, but most often characterized by a smaller size, yellow flowers, and an affinity for rocky habitats. The subspecies may be polyphyletic.
Dudleya attenuata is a species of perennial succulent plant known by the common name taper-tip liveforever, native to Baja California and a small portion of California. It is a rosette-forming leaf succulent which has narrow pencil-shaped leaves that can often be found covered in a white epicuticular wax. The thin, sprawling stems branch to form the clusters of rosettes, with plants creating a "clump" up to 40 cm wide. The small flowers are white or yellow, with 5 spreading petals. It is a diverse, variable species that extends from the southernmost coast of San Diego County to an area slightly north of the Vizcaino Desert, hybridizing with many other species of Dudleya in its range. Some plants with white or pinkish flowers were referred to as Orcutt's liveforever, referring to a former subspecies split on the basis of the flower color.
Dudleya guadalupensis is a very rare species of succulent perennial plant in the family Crassulaceae commonly known as the Guadalupe liveforever. It is a rosette-forming leaf succulent, with foliage that is variously colored light green, green, and a waxy white. It is characterized by dense leaves that fold over the center in dormancy, a curving, sinuous flower stalk, and white, cup-shaped flowers. It is endemic to the rocks and islets off of Guadalupe Island, an isolated volcanic island in the Pacific Ocean located 241 kilometers off of the coast of Baja California.
Dudleya formosa, known by the common name La Misión liveforever, is a species of perennial succulent plant endemic to the Guadalupe Valley in Baja California. It is characterized by bright green leaves, red floral stems, and pink flowers.
Dudleya cochimiana, commonly known as the Cochimíliveforever, is a species of succulent plant in the family Crassulaceae endemic to Cedros Island, a large island off of the coast of Baja California, Mexico. It is a rosette-forming leaf succulent characterized by broad, green to white leaves, and flowers with white to pink petals. It can be found on rocky slopes and canyons along the island.
Dudleya cymosasubsp. costatifolia, known commonly as the Pierpoint Springs dudleya or the Pierpoint Springs liveforever, is a species of succulent plant in the family Crassulaceae, narrowly endemic to a locality in Tulare County, California, United States. It is a clumping plant with small rosettes and bright yellow flowers, resembling alpine cushion plants.
Dudleya cymosasubsp. pumila, most commonly known as the low canyon dudleya, chalky canyon dudleya or California live-forever, is a species of perennial succulent plant. It has diamond to spoon shaped leaves, sometimes coated with a fine white powder, and in May through July, bright red, orange or yellow flowers adorn the short inflorescence. A leaf succulent primarily found growing in rocky cliffs and slopes, it is endemic to California, and grows in the Transverse Ranges and South Coast Ranges, with some outlying populations. A variable plant, in some localities it is difficult to distinguish from other plants in the genus.
Dudleya saxosasubsp. aloides is a species of perennial succulent plant in the family Crassulaceae known by the common names desert dudleya or desert savior. It is a rosette-forming species widely distributed throughout the Peninsular Ranges and desert mountains of California in the United States. It is characterized by bright-yellow or greenish-yellow flowers, and can be found in shaded crevices and slopes. Plants in western half of the range may grade into Dudleya lanceolata.
Dudleya cymosasubsp. cymosa is a species of succulent perennial plant in the family Crassulaceae native to California. It is the autonymous subspecies for Dudleya cymosa, and is known by the common name canyon liveforever. It is native to the California Coast Ranges, the Sierra Nevada and the Santa Monica Mountains. It is characterized by bright-yellow, orange or red flowers and broad, wide leaves. This plant is commonly found growing on rocky outcrops, talus slopes, and in shaded canyons.
Dudleya saxosasubsp. saxosa is a species of succulent perennial plant endemic only to the western Panamint Range of California. It is known commonly as the Panamint liveforever or Panamint dudleya. It is found on north-facing, granitic or limestone slopes and grows up to 20 cm wide. It is characterized by bright yellow, red tinged flowers, which bloom from May to June.
Dudleya virens subsp. extima is a subspecies of succulent plant in the family Crassulaceae commonly known as the Guadalupe green liveforever. It is a rosette-forming leaf succulent, with both green and white waxy foliage. It has white flowers with spreading petals that bloom from May to June. It is a somewhat small plant, continuing a southward trend of decreasing size relative to other Dudleya virens subspecies. This plant is endemic to Guadalupe Island in the eastern Pacific Ocean, which is 241 kilometers off of the Baja California coast. It is very rare, with this plant only surviving on sheer cliff faces and canyons, out of the reach of the former feral goat population rampant on the island. It closely resembles a miniature version of Dudleya virens subsp. virens, but it may be more nearly related to the local Dudleya guadalupensis.
