Equilabium

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Equilabium
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Asterids
Order: Lamiales
Family: Lamiaceae
Tribe: Ocimeae
Genus: Equilabium
Mwany., A.J.Paton & Culham [1] [2]

Equilabium is a genus of flowering plants in the family Lamiaceae. It was split off from the genus Plectranthus in 2018 as the result of a molecular phylogenetic study. Most species are native to Africa, with two found in the Indian subcontinent. [2]

Contents

Description

Species of Equilabium are herbaceous or soft-wooded shrubs, rarely woody shrubs. The herbaceous species may be annual or perennial. The leaves are opposite. The inflorescences are "thryses" – compound structures in which the flowers are arranged on secondary branches. Individual flowers have stalks (pedicels). The sepals form a two-lipped funnel shape, the upper lip having four lobes, the lower lip one lobe. The petals form a two-lipped tube, with an S-shaped basal portion. There are four stamens, whose filaments are not fused together. The style is divided into two parts (bifid). The nutlets are ovoid. [2]

Equilabium and Plectranthus species are distinguished from Coleus by having the stem (pedicel) of the calyx attached symmetrically to the base of the calyx tube, rather than opposite the upper lip, and having the corolla lobes more or less equal in length. Equilabium species can be distinguished from Plectranthus by the truncated shape of the throat of the calyx and by the usually S-shaped tube of the corolla, which is parallel-sided at the base. [2]

Taxonomy

The genus Equilabium was first described in 2018 as the result of a molecular phylogenetic study. Equilabium was split off from the genus Plectranthus , which was discovered not to be monophyletic. Only the names of the genus and the type species, Equilabium laxiflorum , were formally published at the time. [3] The names of 41 more species were published in 2019. [2]

Phylogeny

In 2019, Paton et al. published a summary cladogram for the subtribe Plectranthinae, [2] based on an earlier 2018 study. [3] The new genus Equilabium was established and Coleus revived. [2] In the version below, the three genera accepted in these studies that were formerly included in a broad circumscription of Plectranthus are highlighted.

Tribe Ocimeae

Subtribe Ociminae

Subtribe Plectranthinae

Alvesia (3 spp.)

Aeollanthus (42 spp.)

Tetradenia + Thorncroftia (29 spp.)

Plectranthus (72 spp.)

Capitanopsis (6 spp.)

Equilabium (42 spp.)

Coleus (294 spp.)

Species

Paton et al. (2019) list 42 species: [2] [4]

Related Research Articles

<i>Solenostemon</i> Genus of plants

Solenostemon is a former genus of flowering plants in the family Lamiaceae. It has been included in the genus Plectranthus, but is now included in an expanded Coleus. They are native to tropical Africa, Asia and Australia. Some species formerly placed in this genus are cultivated for their highly variegated leaves.

<i>Ocimum</i> Genus of flowering plants

Ocimum is a genus of aromatic annual and perennial herbs and shrubs in the family Lamiaceae, native to the tropical and warm temperate regions of all 6 inhabited continents, with the greatest number of species in Africa. It is the genus of basil and its best known species are the cooking herb great basil, O. basilicum, and the medicinal herb tulsi, O. tenuiflorum.

<i>Plectranthus</i> Family of shrubs

Plectranthus is a genus of about 85 species of flowering plants from the sage family, Lamiaceae, found mostly in southern and tropical Africa and Madagascar. Common names include spur-flower. Plectranthus species are herbaceous perennial plants, rarely annuals or soft-wooded shrubs, sometimes succulent; sometimes with a tuberous base.

<i>Orthosiphon</i> Genus of flowering plants

Orthosiphon is a genus of plants in the family Lamiaceae native to Africa, Southern Asia and Queensland, with one species (O. americanus) in Colombia. They are herbaceous shrubs which grow to a height of 1.5 m (5 ft). Some Orthosiphon species are popular garden plants because of their flowers, which are white and bluish with filaments resembling a cat's whiskers. In the wild, the plants can be seen growing in forests and along roadsides.

<i>Coleus caninus</i> Species of flowering plant

Coleus caninus, synonym Plectranthus caninus, is a herb from the mint family Lamiaceae, native to southern and eastern Africa from Angola to Sudan and to India and Myanmar.

<i>Coleus scutellarioides</i> Species of flowering plant

Coleus scutellarioides, commonly known as coleus, is a species of flowering plant in the family Lamiaceae, native to southeast Asia through to Australia. Typically growing to 60–75 cm (24–30 in) tall and wide, it is a bushy, woody-based evergreen perennial, widely grown for the highly decorative variegated leaves found in cultivated varieties. Another common name is painted nettle, reflecting its relationship to deadnettles, which are in the same family. The synonyms Coleus blumei, Plectranthus scutellarioides and Solenostemon scutellarioides are also widely used for this species.

<i>Coleus</i> Genus of flowering plants

Coleus is a genus of annual or perennial herbs or shrubs, sometimes succulent, sometimes with a fleshy or tuberous rootstock, found in the Afro-Eurasia tropics and subtropics.

<i>Tinnea</i> Genus of flowering plants

Tinnea (sunbells) is a genus of plants in the family Lamiaceae first described in 1867. It is native to sub-Saharan Africa. It was named in honour of the Dutch explorer Alexine Tinne.

  1. Tinnea aethiopicaKotschy ex Hook.f. - widespread from Mali to Somalia south to Mozambique; naturalized in Trinidad & Tobago
  2. Tinnea antiscorbuticaWelw. - DRC, Zambia, Angola
  3. Tinnea apiculataRobyns & Lebrun - eastern Africa from Rwanda to Mozambique
  4. Tinnea barbataVollesen - Eswatini, northern South Africa
  5. Tinnea barteriGürke - western Africa
  6. Tinnea benguellensisGürke - Angola
  7. Tinnea coeruleaGürke - DRC, Zambia, Angola
  8. Tinnea eriocalyxWelw. - DRC, Angola, Botswana, Namibia
  9. Tinnea galpiniiBriq. - Eswatini, Mozambique, South Africa
  10. Tinnea gossweileriRobyns & Lebrun - Angola
  11. Tinnea gracilisGürke - Tanzania to Zambia
  12. Tinnea mirabilis(Bullock) Vollesen - Tanzania
  13. Tinnea physalisE.A.Bruce - Tanzania
  14. Tinnea platyphyllaBriq. - DRC
  15. Tinnea rhodesianaS.Moore - South Africa, Namibia, Zimbabwe, Zambia, Angola, Mozambique
  16. Tinnea somalensisGürke ex Chiov. - Ethiopia
  17. Tinnea vesiculosaGürke - Tanzania, Malawi
  18. Tinnea vestitaBaker - Zimbabwe, Zambia, Angola, Botswana
  19. Tinnea zambesiacaBaker - Zimbabwe, Zambia, Malawi, Mozambique
<i>Aeollanthus</i> Genus of flowering plants

Aeollanthus (rocksage) is a genus in the mint family, Lamiaceae. All the species are native to Africa.

<i>Capitanopsis</i> Genus of flowering plants

Capitanopsis is a genus of plants in the family Lamiaceae, first described in 1916. It contains six known species, all endemic to Madagascar.

Endostemon (keepsafe) is a genus of plants in the family Lamiaceae, first described in 1910. It is native primarily to eastern Africa, with some species in central and southern Africa, the Arabian Peninsula, Madagascar, and the Indian subcontinent.

  1. Endostemon albusA.J.Paton, Harley & M.M.Harley - Kenya, Tanzania, Mozambique
  2. Endostemon camporum(Gürke) M.R.Ashby - Kenya, Tanzania
  3. Endostemon ctenoneurusHarley - Kenya, Somalia
  4. Endostemon glandulosusHarley & Sebsebe - Ethiopia
  5. Endostemon gracilis(Benth.) M.R.Ashby - Kenya, Somalia, Tanzania, Yemen
  6. Endostemon kelleri(Briq.) Ryding ex A.J.Paton & Harley - Kenya, Somalia, Ethiopia
  7. Endostemon leucosphaerus(Briq.) A.J.Paton, Harley & M.M.Harley - Somalia, Ethiopia
  8. Endostemon membranaceus(Benth.) Ayob. ex A.J.Paton & Harley - Cameroon, Angola, Central African Republic
  9. Endostemon obbiadensis(Chiov.) M.R.Ashby - Somalia
  10. Endostemon obtusifolius(E.Mey.) N.E.Br. - from South Africa north to Angola and Tanzania
  11. Endostemon racemosusRyding, A.J.Paton & Thulin - Somalia
  12. Endostemon stenocaulis(Hedge) Ryding, A.J.Paton & Thulin - Somalia
  13. Endostemon tenuiflorus(Benth.) M.R.Ashby - eastern + southern Africa, Madagascar, Arabian Peninsula
  14. Endostemon tereticaulis(Poir.) M.R.Ashby - widespread across much of tropical Africa, also Yemen + Saudi Arabia
  15. Endostemon tomentosusHarley & Sebsebe - Somalia
  16. Endostemon tubulascens(Briq.) M.R.Ashby - Angola
  17. Endostemon usambarensisM.R.Ashby - Tanzania
  18. Endostemon villosus(Briq.) M.R.Ashby - central Africa
  19. Endostemon viscosus(Roth) M.R.Ashby - India, Assam, Sri Lanka
  20. Endostemon wakefieldii(Baker) M.R.Ashby - Kenya
<i>Haumaniastrum</i> Genus of flowering plants

Haumaniastrum is a genus of flowering plants in the mint family, Lamiaceae, first described in 1959. The species are native to Africa.

<i>Pycnostachys</i> Genus of flowering plants

Pycnostachys is a genus of plants in the family Lamiaceae, first described in 1826. It is native to sub-Saharan Africa including Madagascar.

  1. Pycnostachys abyssinicaFresen. - Ethiopia
  2. Pycnostachys angolensisG.Taylor - Angola
  3. Pycnostachys batesiiBaker - Cameroon, Zaïre, Sudan, Uganda
  4. Pycnostachys chevalieriBriq. - Central African Republic
  5. Pycnostachys ciliataBramley - Tanzania, Malawi, Mozambique, Zambia
  6. Pycnostachys coeruleaHook. - East Africa from Ethiopia to Botswana; Madagascar
  7. Pycnostachys congensisGürke - Zaïre, Zambia
  8. Pycnostachys de-wildemanianaRobyns & Lebrun - East Africa from Burundi to Zimbabwe
  9. Pycnostachys deflexifoliaBaker - Kenya, Tanzania, Uganda
  10. Pycnostachys descampsiiBriq. - Zaïre
  11. Pycnostachys elliotiiS.Moore - Zaïre, Uganda
  12. Pycnostachys eminiiGürke - from Cameroon to Ethiopia + Tanzania
  13. Pycnostachys erici-roseniiR.E.Fr - Zaïre, Uganda, Burundi, Rwanda
  14. Pycnostachys goetzeniiGürke - Zaïre, Uganda, Rwanda
  15. Pycnostachys gracilisR.D.Good - Angola
  16. Pycnostachys graminifoliaPerkins - Tanzania
  17. Pycnostachys kassneriDe Wild - Zaïre, Zambia, Tanzania
  18. Pycnostachys lancifoliaBramley - Tanzania
  19. Pycnostachys meyeriGürke - western + central Africa
  20. Pycnostachys nepetifoliaBaker - Kenya
  21. Pycnostachys niamniamensisGürke - South Sudan, Kenya, Uganda
  22. Pycnostachys orthodontaGürke - Zimbabwe, Mozambique, Zambia, Tanzania
  23. Pycnostachys parvifoliaBaker - Zambia, Tanzania, Malawi
  24. Pycnostachys prittwitziiPerkins - Zambia, Tanzania
  25. Pycnostachys pseudospeciosaBuscal. & Muschl. - Zaïre, Zambia
  26. Pycnostachys recurvataRyding - Ethiopia
  27. Pycnostachys reticulata(E.Mey.) Benth. - central + southern Africa
  28. Pycnostachys ruandensisDe Wild. - east-central Africa
  29. Pycnostachys schliebeniiMildbr. - east-central Africa
  30. Pycnostachys schweinfurthiiBriq - western + central Africa
  31. Pycnostachys speciosaGürke - Kenya, Uganda, Tanzania, Rwanda
  32. Pycnostachys sphaerocephalaBaker - Zambia, Tanzania, Malawi, Zaïre
  33. Pycnostachys stuhlmanniiGürke - central Africa
  34. Pycnostachys umbrosa(Vatke) Perkins - Kenya, Tanzania
  35. Pycnostachys urticifoliaHook. - southeastern Africa
  36. Pycnostachys verticillataBaker - Zambia, Tanzania
<i>Syncolostemon</i> Genus of flowering plants

Syncolostemon (sagebushes) is a genus of plants in the family Lamiaceae, first described in 1838. It is native primarily to South Africa, with some species in other parts of sub-Saharan Africa, plus one species in India.

  1. Syncolostemon albiflorus(N.E.Br.) D.F.Otieno - Transvaal, Eswatini
  2. Syncolostemon argenteusN.E.Br. - KwaZulu-Natal
  3. Syncolostemon bolusii(N.E.Br.) D.F.Otieno - KwaZulu-Natal
  4. Syncolostemon bracteosus(Benth.) D.F.Otieno - widespread across much of sub-Saharan Africa
  5. Syncolostemon canescens(Gürke) D.F.Otieno - Angola, Botswana, Zimbabwe, Eswatini, South Africa
  6. Syncolostemon cinereum(Codd) D.F.Otieno & Retief - South Africa
  7. Syncolostemon comosus(Wight ex Benth.) D.F.Otieno - southern India
  8. Syncolostemon comptoniiCodd - Eswatini
  9. Syncolostemon concinnusN.E.Br. - Eswatini, South Africa
  10. Syncolostemon densiflorusBenth. - South Africa
  11. Syncolostemon elliottii(Baker) D.F.Otieno - Zimbabwe, Botswana, Transvaal
  12. Syncolostemon eriocephalusVerd. - Northern Province of South Africa
  13. Syncolostemon flabellifolius(S.Moore) A.J.Paton - Chimanimani Mountains of Mozambique + Zimbabwe
  14. Syncolostemon floccosus(Launert) D.F.Otieno - Namibia
  15. Syncolostemon foliosus(S.Moore) D.F.Otieno - Eswatini, South Africa
  16. Syncolostemon gerrardii(N.E.Br.) D.F.Otieno - South Africa
  17. Syncolostemon incanus(Codd) D.F.Otieno - Northern Province of South Africa
  18. Syncolostemon latidens(N.E.Br.) Codd - KwaZulu-Natal
  19. Syncolostemon linearis(Benth.) D.F.Otieno - Zimbabwe
  20. Syncolostemon macranthus(Gürke) Ashby - Drakensberg Mountains in South Africa
  21. Syncolostemon macrophyllusGürke - South Africa
  22. Syncolostemon madagascariensis(A.J.Paton & Hedge) D.F.Otieno - Madagascar
  23. Syncolostemon modestus(Codd) D.F.Otieno - Eswatini, South Africa
  24. Syncolostemon namapaensisD.F.Otieno - Mozambique, Tanzania
  25. Syncolostemon obermeyerae(M.Ashby) D.F.Otieno - Northern Province of South Africa
  26. Syncolostemon oritrephes(Wild) D.F.Otieno - Chimanimani Mountains of Mozambique + Zimbabwe
  27. Syncolostemon ornatus(S.Moore) D.F.Otieno - Chimanimani Mountains of Zimbabwe
  28. Syncolostemon parviflorusE.Mey. ex Benth. - Eswatini, South Africa
  29. Syncolostemon parvifolius(Codd) D.F.Otieno - Northern Province of South Africa
  30. Syncolostemon persimilis(N.E.Br.) D.F.Otieno - Northern Province of South Africa
  31. Syncolostemon petiolatus(Ashby) D.F.Otieno - Eswatini, South Africa, Mozambique
  32. Syncolostemon pretoriae(Gürke) D.F.Otieno - Eswatini, South Africa
  33. Syncolostemon punctatus(Codd) D.F.Otieno - Northern Province of South Africa
  34. Syncolostemon ramosus(Codd) D.F.Otieno - KwaZulu-Natal
  35. Syncolostemon ramulosusE.Mey. ex Benth. - KwaZulu-Natal, Cape Province
  36. Syncolostemon rehmannii(Gürke) D.F.Otieno - Northern Province of South Africa
  37. Syncolostemon rotundifoliusE.Mey. ex Benth. - KwaZulu-Natal, Cape Province
  38. Syncolostemon rugosifolius(M.Ashby) D.F.Otieno - Northern Province of South Africa
  39. Syncolostemon stalmansii(A.J.Paton & K.Balkwill) D.F.Otieno - Mpumalanga, Eswatini
  40. Syncolostemon stenophyllus(Gürke) D.F.Otieno - KwaZulu-Natal, Cape Province
  41. Syncolostemon subvelutinus(Gürke) D.F.Otieno - Northern Province of South Africa
  42. Syncolostemon teucriifolius(Hochst.) D.F.Otieno - Mozambique, Zimbabwe, South Africa
  43. Syncolostemon thorncroftii(N.E.Br.) D.F.Otieno - Northern Province of South Africa
  44. Syncolostemon transvaalensis(Schltr.) D.F.Otieno - Northern Province of South Africa
  45. Syncolostemon welwitschii(Rolfe) D.F.Otieno - Nigeria, Cameroon, Central African Republic, Zaire, Tanzania, Zambia, Angola

Coleus maculosus is a species of flowering plant in the family Lamiaceae. It is widespread in tropical Africa and Madagascar.

<i>Coleus comosus</i> Species of flowering plant

Coleus comosus, synonym Plectranthus ornatus, is a flowering plant from the mint family Lamiaceae, native to eastern Africa.

<i>Capitanopsis brevilabra</i> Species of flowering plant

Capitanopsis brevilabra is a species of flowering plant in the family Lamiaceae. It is an herb or shrub native to southeastern Madagascar. It flowers in March.

Capitanopsis oreophila, synonym Plectranthus bipinnatus, is a species of flowering plant in the family Lamiaceae. It is a small shrub native to Madagascar, where it is commonly known as andriamborondrao.

References

  1. "Equilabium Mwany., A.J.Paton & Culham", The International Plant Names Index , retrieved 2020-07-29
  2. 1 2 3 4 5 6 7 8 Paton, Alan J.; Mwanyambo, Montfort; Govaerts, Rafaël H.A.; Smitha, Kokkaraniyil; Suddee, Somran; Phillipson, Peter B.; Wilson, Trevor C.; Forster, Paul I. & Culham, Alastair (2019), "Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera", PhytoKeys (129): 1–158, doi: 10.3897/phytokeys.129.34988 , PMC   6717120 , PMID   31523157
  3. 1 2 Paton, A.; Mwanyambo, M. & Culham, A. (2018). "Phylogenetic study of Plectranthus, Coleus and allies (Lamiaceae): Taxonomy, distribution and medicinal use" (PDF). Botanical Journal of the Linnean Society. 188 (4): 355–376. doi: 10.1093/botlinnean/boy064 .
  4. "Equilabium Mwany., A.J. Paton & Culham". Tropicos . Missouri Botanical Garden . Retrieved 2020-07-29.