Equus ovodovi Temporal range: Late Pleistocene-Holocene | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Perissodactyla |
Family: | Equidae |
Genus: | Equus |
Species: | †E. ovodovi |
Binomial name | |
†Equus ovodovi Eisenmann and Vasiliev, 2011 | |
Equus ovodovi is an extinct species of equine known from the Late Pleistocene and Holocene of North and East Asia.
Equus ovodovi was slightly larger than the Asiatic wild ass, [1] from which it can be distinguished based on its larger and more robust limb bones. [2]
Isotopic analysis of specimens from the Holocene of China suggests a preference for C4 plants. [3]
Equus ovodovi has been suggested to be the last surviving member of the subgenus Sussemionus, which first appeared in North America over 2 million years ago, and was formerly present across Afro-Eurasia. [1] [4] Some later studies questioned its relationship to the subgenus Sussemionus. [5] Remains are known spanning from southern Western Siberia to Northern China, with the youngest remains dating to around 3500 years ago (~1500 BC) in Northern China. Genetic evidence suggests that it more closely related to zebras and asses than to horses. Initial genetic analysis based on the mitochondrial DNA found the exact relationship to be uncertain, but analysis of the full nuclear genome suggests that zebras and asses are more closely related to each other than either are to E. ovodovi, though there had been gene flow into the Equus ovodovi lineage from both the last common ancesor of zebras and asses and the last common ancestor of zebras, as well as gene flow from the Equus ovodovi lineage into the last common ancestor of asses. [6]
Cladogram after Cai et al. 2022: [6]
Equus |
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Its genetic diversity progressively declined to very low levels over the course of the Late Pleistocene and Holocene prior to its extinction. [6]
Equidae is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. The family evolved around 50 million years ago from a small, multi-toed ungulate into larger, single-toed animals. All extant species are in the genus Equus, which originated in North America. Equidae belongs to the order Perissodactyla, which includes the extant tapirs and rhinoceros, and several extinct families.
Perissodactyla is an order of ungulates. The order includes about 17 living species divided into three families: Equidae, Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). They typically have reduced the weight-bearing toes to three or one of the five original toes, though tapirs retain four toes on their front feet. The nonweight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, artiodactyls bear most of their weight equally on four or two of the five toes: their third and fourth toes. Another difference between the two is that perissodactyls digest plant cellulose in their intestines, rather than in one or more stomach chambers as artiodactyls, with the exception of Suina, do.
The quagga is an extinct subspecies of the plains zebra that was endemic to South Africa until it was hunted to extinction in the late 19th century. It was long thought to be a distinct species, but early genetic studies have supported it being a subspecies of plains zebra. A more recent study suggested that it was the southernmost cline or ecotype of the species.
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The onager, also known as hemione or Asiatic wild ass, is a species of the family Equidae native to Asia. A member of the subgenus Asinus, the onager was described and given its binomial name by German zoologist Peter Simon Pallas in 1775. Six subspecies have been recognized, two of which are extinct.
The woolly rhinoceros, simply known as woolly rhino, is an extinct species of rhinoceros that inhabited northern Eurasia during the Pleistocene epoch. The woolly rhinoceros was a member of the Pleistocene megafauna. The woolly rhinoceros was covered with long, thick hair that allowed it to survive in the extremely cold, harsh mammoth steppe. It had a massive hump reaching from its shoulder and fed mainly on herbaceous plants that grew in the steppe. Mummified carcasses preserved in permafrost and many bone remains of woolly rhinoceroses have been found. Images of woolly rhinoceroses are found among cave paintings in Europe and Asia. The species range contracted towards Siberia beginning around 17,000 years ago, with the youngest known records being around 14,000 years old in northeast Siberia, coinciding with the Bølling–Allerød warming, which likely disrupted its habitat, with environmental DNA records possibly extending the range of the species around 9,800 years ago.
Equus is a genus of mammals in the family Equidae, which includes horses, asses, and zebras. Within the Equidae, Equus is the only recognized extant genus, comprising seven living species. Like Equidae more broadly, Equus has numerous extinct species known only from fossils. The genus most likely originated in North America and spread quickly to the Old World. Equines are odd-toed ungulates with slender legs, long heads, relatively long necks, manes, and long tails. All species are herbivorous, and mostly grazers, with simpler digestive systems than ruminants but able to subsist on lower-quality vegetation.
The kiang is the largest of the Asinus subgenus. It is native to the Tibetan Plateau in Ladakh India, northern Pakistan, Tajikistan, China and northern Nepal. It inhabits montane grasslands and shrublands. Other common names for this species include Tibetan wild ass, khyang and gorkhar.
The wild horse is a species of the genus Equus, which includes as subspecies the modern domesticated horse as well as the endangered Przewalski's horse. The European wild horse, also known as the tarpan, that went extinct in the late 19th or early 20th century has previously been treated as the nominate subspecies of wild horse, Equus ferus ferus, but more recent studies have cast doubt on whether tarpans were truly wild or if they actually were feral horses or hybrids.
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The European wild ass or hydruntine is an extinct equine from the Middle Pleistocene to Late Holocene of Europe and West Asia, and possibly North Africa. It is a member of the subgenus Asinus, and closely related to the living Asiatic wild ass. The specific epithet, hydruntinus, means from Otranto.
Equus sivalensis is an extinct species of large equid native to the northern Indian subcontinent. Remains date from the beginning of the Pleistocene, c. 2.58 million years ago until around 600,000 years ago, during the Middle Pleistocene. It is considered a "stenonine horse", meaning that it is more closely related to zebras and asses than true horses. Based on isotopes and teeth morphology, it is thought to have been a grazer. The later species Equus namadicus from the same region has sometimes been suggested to be a synonym due to their similar teeth morphology.
Hippidion is an extinct genus of equine that lived in South America from the Late Pliocene to the end of the Late Pleistocene (Lujanian), between 2.5 million and 11,000 years ago. They were one of two lineages of equines native to South America during the Pleistocene epoch, alongside Equus (Amerhippus) neogeus.
The evolution of the horse, a mammal of the family Equidae, occurred over a geologic time scale of 50 million years, transforming the small, dog-sized, forest-dwelling Eohippus into the modern horse. Paleozoologists have been able to piece together a more complete outline of the evolutionary lineage of the modern horse than of any other animal. Much of this evolution took place in North America, where horses originated but became extinct about 10,000 years ago, before being reintroduced in the 15th century.
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Stephanorhinus is an extinct genus of two-horned rhinoceros native to Eurasia and North Africa that lived during the Late Pliocene to Late Pleistocene. Species of Stephanorhinus were the predominant and often only species of rhinoceros in much of temperate Eurasia, especially Europe, for most of the Pleistocene. The last two species of Stephanorhinus – Merck's rhinoceros and the narrow-nosed rhinoceros – went extinct during the last glacial period.
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