Fibla carpenteri Temporal range: | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Raphidioptera |
Family: | Inocelliidae |
Genus: | Fibla |
Subgenus: | Fibla |
Species: | †F. carpenteri |
Binomial name | |
†Fibla carpenteri Engel, 1995 | |
Fibla carpenteri is an extinct species of snakefly in the Inocelliidae genus Fibla . [1] F. carpenteri is named in honor of the paleoentomologist Dr Frank Carpenter, for his vast knowledge and interest in Raphidioptera. [1]
The species is known from a single specimen, the holotype, deposited in the Harvard University, Museum of Comparative Zoology as specimen #9999. Dr. Michael S. Engel first studied and described the species after finding the specimen in the Harvard collections. He published his type description in the journal Psyche volume 102 published in 1995. [1] Fairly well preserved in Eocene Baltic amber, the female individual has a torn forewing missing the distal portion, partial antennae, and the ovipositor is severed and missing the tip. There are also a number of small areas with "schimmel", a type of white mold sometimes present on arthropods in amber. [1] With a total length, not including ovipositor or antennae, of just over 18 millimetres (0.71 in), Fibla carpenteri is the largest species of snakefly from amber and the largest species of the genus. [1] As a whole the female shows no light color marking and was a fairly uniform dark brown to black coloration. The wings are hyaline with brown coloration of the vein structure and are slightly fuscous at the base. [1] The pterostigma is also colored brown.
F. carpenteri is one of only four extinct Fibla which are known from the fossil record. [2] Along with F. erigena F. carpenteri is one of two known from the baltic amber deposits, while F. cerdanica is from the Miocene of Spain and F. exusta is from the Eocene of the Florissant Formation, Colorado. [2]
Snakeflies are a group of predatory insects comprising the order Raphidioptera with two extant families: Raphidiidae and Inocelliidae, consisting of roughly 260 species. In the past, the group had a much wider distribution than it does now; snakeflies are found in temperate regions worldwide but are absent from the tropics and the Southern Hemisphere. Recognisable representatives of the group first appeared during the Early Jurassic. They are a relict group, having reached their apex of diversity during the Cretaceous before undergoing substantial decline.
Frank Morton Carpenter was an American entomologist and paleontologist. He received his PhD from Harvard University, and was curator of fossil insects at the Harvard Museum of Comparative Zoology for 60 years. He studied the Permian fossil insects of Elmo, Kansas, and compared the North American fossil insect fauna with Paleozoic taxa known from elsewhere in the world. A careful and methodical worker, he used venation and mouthparts to determine the relationships of fossil taxa, and was author of the Treatise volume on Insects. He reduced the number of extinct insect orders then described from about fifty to nine.
The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that these forests created more than 100,000 tons of amber. Today, more than 90% of the world's amber comes from Kaliningrad Oblast of Russia. It is a major source of income for the region; the local Kaliningrad Amber Combine extracted 250 tonnes of it in 2014, 400 tonnes in 2015.
Inocelliidae is a small family of snakeflies containing 8 genera of which one is known only from fossils. They are commonly known as inocelliid snakeflies. The largest known species is Fibla carpenteri known from fossils found in baltic amber.
Nanoraphidia is an extinct genus of snakefly in the family Mesoraphidiidae containing the species Nanoraphidia electroburmica and Nanoraphidia lithographica.
Electrinocellia is an extinct monotypic genus of snakefly in the family Inocelliidae containing the single species Electrinocellia peculiaris and known from Eocene aged Baltic amber.
Florissantoraphidia is an extinct genus of snakefly in the family Raphidiidae containing two described species Florissantoraphidia funerata and Florissantoraphidia mortua. Both species were originally placed in the living raphidiid genus Raphidia, as Raphidia funerata and Raphidia mortua respectively. before being redescribed and transferred to the newly erected genus in 2014.
Ororaphidia is an extinct genus of snakefly containing two species: the type species Ororaphidia megalocephala and Ororaphidia bifurcata.
Styporaphidia is a genus of snakefly, belonging to the extinct family Mesoraphidiidae, containing up to three species, the type species Styporaphidia magia, Styporaphidia willmanni and tentatively Styporaphidia? hispanica. The genus was named from the Greek stypos meaning "stem" or "stump" and Raphidia, the type genus for, and most often used as, a stem for generic names in the order Raphidioptera. The species name of S. magia is from the Greek word mageia meaning "magic" while the species name for S.? hispanica is from the Latin Hispania meaning "Spain" in reference to the type locality of the species.
Archiinocellia is an extinct genus of snakefly in the family Raphidiidae known from Eocene fossils found in western North America. The genus contains two species, the older Archiinocellia oligoneura and the younger Archiinocellia protomaculata. The type species is of Ypresian age and from the Horsefly Shales of British Columbia, while the younger species from the Lutetian Green River Formation in Colorado. Archiinocellia protomaculata was first described as Agulla protomaculata, and later moved to Archiinocellia.
Proraphidia is a genus of snakefly in the extinct family Mesoraphidiidae. The genus currently contains three species; Proraphidia gomezi from the La Pedrera de Rúbies Formation in Spain, Proraphidia hopkinsi from the Weald Clay in England, and the type species Proraphidia turkestanica from Kazakhstan. The genus was first described by O. M. Martynova in 1941 with the publication of P. turkestanica from Jurassic deposits of the Karabastau Formation in Karatau, Kazakhstan.
Neanaperiallus is an extinct monotypic genus of parasitic wasp in the Eupelmidae subfamily Neanastatinae containing only a single species, Neanaperiallus masneri. The genus is solely known from the Early Eocene Baltic amber deposits in the Baltic Sea region of Europe.
Kachinus is an extinct genus of ant-like stone beetle in the family Staphylinidae containing the single species Kachinus antennatus.
Agulla mineralensis is an extinct species of snakefly in the raphidiid genus Agulla. The species is solely known from the Middle Miocene, late Bartovian stage, Pacific Union Site in the Stewart Valley Group, Mineral County, Nevada.
Amarantoraphidia is an extinct genus of snakefly in the family Mesoraphidiidae. The genus is solely known from Early Cretaceous, Albian age, fossil amber found in Spain. Currently the genus comprises only a single species Amarantoraphidia ventolina.
Alavaraphidia is an extinct genus of snakefly in the family Mesoraphidiidae. The genus is solely known from an Early Cretaceous, Albian age, fossil amber found in Spain. Currently, the genus comprises a single species, Alavaraphidia imperterrita.
Mesoraphidiidae is an extinct family of snakeflies in the suborder Raphidiomorpha. The family lived from the Late Jurassic through the Late Cretaceous and is known from twenty-five genera. Mesoraphidiids have been found as both compression fossils and as inclusions in amber. The family was first proposed in 1925 by the Russian paleoentomologist Andrey Vasilyevich Martynov based on Upper Jurassic fossils recovered in Kazakhstan. The family was expanded in 2002 by the synonymizing of several other proposed snakefly families. The family was divided into three subfamilies and one tribe in a 2011 paper, further clarifying the relationships of the included genera.
Lebanoraphidia is an extinct genus of snakefly in the family Mesoraphidiidae. The genus is solely known from Cretaceous, Upper Neocomian, fossil amber found in Lebanon. Currently the genus is composed of a single species Lebanoraphidia nana.
Electrostephanus is an extinct genus of crown wasp in the hymenopteran family Stephanidae, and is the only genus placed in the subfamily Electrostephaninae. The genus contains four described species, E. brevicornis, E. neovenatus, E. janzeni, and E. petiolatus, placed in two subgenera E. (Electrostephanus) and E. (Electrostephanodes). Electrostephanus is known from several middle Eocene fossils which have been found in Europe.
Dicromantispa moronei is an extinct species of mantidfly in the neuropteran family Mantispidae known from a fossil found in the Caribbean.