Fordilloidea

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Fordilloidea
Temporal range: 520.0–513.0  Ma
Fordilla troyensis shell.jpg
Fordilla troyensis outer shell surface [1]
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Mollusca
Class: Bivalvia
Order: Fordillida
Superfamily: Fordilloidea
Pojeta, 1975
Families

Fordilloidea is an extinct superfamily of early bivalves [2] containing two described families, Fordillidae and Camyidae and the only superfamily in the order Fordillida. [2] The superfamily is known from fossils of early to middle Cambrian age found in North America, Greenland, Europe, the Middle East, Asia, and Australia. [3] [4] [5] Fordillidae currently contains two genera, Fordilla and Pojetaia each with up to three described species while Camyidae only contains a single genus Camya with one described species, Camya asy. [3] [5] Due to the size and age of the fossil specimens, Fordillidae species are included as part of the Turkish Small shelly fauna. [3]

Description

Fordilla are small bivalves with valves that are equal in size and suboval in shape. In size, Fordilla specimens reach a total shell length of up to 4 millimetres (0.16 in) and a height of 25 millimetres (0.98 in). [3] [6] The shells are compressed laterally and the back edge is slightly broadened. The rear adductor is less developed and smaller than the front adductor, while the small pedal retractor muscle scar is positioned near the front adductor scar. [3] The valve hinge is usually straight to slightly convexly curved and each valve will have at most one tooth present. The external surface of the shell occasionally shows faint ribbing. [3]

Similar to Fordilla, species of Pojetaia are small, with valves to less than 2 millimetres (0.079 in) in length. Pojetaia species have an overall shape which is suboval, with the subequal valves slightly elongated. The ligament is straight with an umbo which is central to subcentral. In contrast to Fordilla, the rear adductor muscle was larger and more developed than the front adductor, with pallial muscles arranged along the valve margins. Also in contrast to Fordilla, valves of Pojetaia possess between one and three teeth, with up to two teeth per valve. The exteriors of the shells show faint ribbing and fine comarginal growth lines. [3]

Camya is based on the fossils of two juvenile specimens which are both incomplete due to only the left valve of each being recovered. The valves have a distinct subtriangular shape and possess a long straight hinge. The umbo is positioned notably anterior on the shell and the beak is bracketed by two teeth of indistinctly pyramidal shape. [3] The presence of the teeth was later questioned in a 1998 study by G. Geyer and M. Streng and cited the lack of preserved muscle scars as reason to suspect the placement of Camya asy in Bivalvia. [3]

The inner shell layers of Fordilla and Pojetaia species both consist of layers of carbonate which is akin to the laminar aragonite layer found in extant monoplacophora. [7] The structuring is similar to shell layering found in the extinct genera Anabarella and Watsonella which is thought to suggest members of the phylum Mollusca developed nacre independently several times. [7] Of the four accepted bivalve genera to have been described from the Cambrian, Fordilloidea contains three, Camya, Fordilla, and Pojetaia , [3] with the remaining genus Tuarangia in the possibly related order Tuarangiida. [2] [8]

Related Research Articles

<span class="mw-page-title-main">Bivalvia</span> Class of molluscs

Bivalvia or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of aquatic molluscs that have laterally compressed soft bodies enclosed by a calcified exoskeleton consisting of a hinged pair of half-shells known as valves. As a group, bivalves have no head and lack some typical molluscan organs such as the radula and the odontophore. Their gills have evolved into ctenidia, specialised organs for feeding and breathing.

<span class="mw-page-title-main">Scallop</span> Common name for several shellfish, many edible

Scallop is a common name that encompasses various species of marine bivalve mollusks in the taxonomic family Pectinidae, the scallops. However, the common name "scallop" is also sometimes applied to species in other closely related families within the superfamily Pectinoidea, which also includes the thorny oysters.

<span class="mw-page-title-main">Venerida</span> Order of molluscs

Venerida is an order of mostly saltwater but also some freshwater bivalve molluscs. This order includes many familiar groups such as many clams that are valued for food and a number of freshwater bivalves.

<span class="mw-page-title-main">Pinnidae</span> Family of bivalves

The Pinnidae are a taxonomic family of large saltwater clams sometimes known as pen shells. They are marine bivalve molluscs in the order Pteriida.

<span class="mw-page-title-main">Bivalve shell</span> Seashell

A bivalve shell is the enveloping exoskeleton or shell of a bivalve mollusc, composed of two hinged halves or valves. The two half-shells, called the "right valve" and "left valve", are joined by a ligament and usually articulate with one another using structures known as "teeth" which are situated along the hinge line. In many bivalve shells, the two valves are symmetrical along the hinge line — when truly symmetrical, such an animal is said to be equivalved; if the valves vary from each other in size or shape, inequivalved. If symmetrical front-to-back, the valves are said to be equilateral, and are otherwise considered inequilateral.

Anabarella is a species of bilaterally-flattened monoplacophoran mollusc, with a morphological similarity to the rostroconchs. Its shell preserves evidence of three mineralogical textures on its outer surface: it is polygonal near the crest of the shell, subsequently changing to both spiny and stepwise. Its internal microstructure is calcitic and semi-nacreous. Its name reflects its provenance from Anabar, Siberia. It has been interpreted as ancestral to the rostroconchs, and has been aligned to the Helcionellidae.

<i>Fordilla</i> Extinct genus of bivalves

Fordilla is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early Cambrian fossils found in North America, Greenland, Europe, the Middle East, and Asia. The genus currently contains three described species, Fordilla germanica, Fordilla sibirica, and the type species Fordilla troyensis.

<span class="mw-page-title-main">Rhynchonelliformea</span> Subphylum of brachiopods

Rhynchonelliformea is a major subphylum and clade of brachiopods. It is roughly equivalent to the former class Articulata, which was used previously in brachiopod taxonomy up until the 1990s. These so-called articulated brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations and separate sets of simple opening and closing muscles.

Pojetaia is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early to Middle Cambrian fossils found in North America, Greenland, Europe, North Africa, Asia, and Australia. The genus currently contains two accepted species, Pojetaia runnegari, the type species, and Pojetaia sarhroensis, though up to seven species have been proposed. The genera Buluniella, Jellia, and Oryzoconcha are all considered synonyms of Pojetaia.

<i>Tellimya ferruginosa</i> Species of bivalve

Tellimya ferruginosa is a species of small marine bivalve mollusc in the family Lasaeidae. It is found on the eastern side of the Atlantic Ocean.

<span class="mw-page-title-main">Fordillidae</span> Extinct family of bivalves

Fordillidae is an extinct family of early bivalves and one of two families in the extinct superfamily Fordilloidea. The family is known from fossils of early to middle Cambrian age found in North America, Greenland, Europe, the Middle East, Asia, and Australia. The family currently contains two genera, Fordilla and Pojetaia, each with up to three described species. Due to the size and age of the fossil specimens, Fordillidae species are included as part of the Turkish Small shelly fauna.

Camya is an extinct genus of early bivalve and is the only genus in the extinct family Camyidae. The genus is known solely from early Middle Cambrian fossils found in Europe. The genus currently contains a solitary accepted species, Camya asy.

Tuarangia is a Cambrian shelly fossil interpreted as an early bivalve, though alternative classifications have been proposed and its systematic position remains controversial. It is the only genus in the extinct family Tuarangiidae and order Tuarangiida. The genus is known solely from Middle to Late Cambrian fossils found in Europe and New Zealand. The genus currently contains two accepted species, Tuarangia gravgaerdensis and the type species Tuarangia paparua.

Concavodonta is an extinct genus of early bivalve in the extinct family Praenuculidae. The genus is one of three genera in the subfamily Concavodontinae. Concavodonta is known solely from late Ordovician, Caradoc epoch, fossils found in Europe and South America. The genus currently contains three accepted species, Concavodonta imbricata, Concavodonta ovalis and the type species Concavodonta ponderata.

Hemiconcavodonta is an extinct genus of bivalve in the extinct family Praenuculidae. The genus is one of three genera in the subfamily Concavodontinae. Hemiconcavodonta is known solely from late Ordovician, Caradoc epoch, fossils found in South America. The genus currently contains a single accepted species, Hemiconcavodonta minuta.

Praenuculinae is an extinct subfamily of prehistoric bivalves in the family Praenuculidae. Praenuculinae species lived from the middle Ordovician through the late Devonian. Praenuculinae fossils are found in Europe, Africa, North America and South America, and species are thought to have been stationary attached to substrate in shallow infaunal marine water environments where they formed shells of an aragonite composition. The subfamily Praenuculinae was named by Teresa M. Sánchez in 1999.

<i>Cucullaea labiata</i> Species of bivalve

Cucullaea labiata is a species of saltwater clam or ark shell, a marine bivalve mollusk in the family Cucullaeidae.

<span class="mw-page-title-main">Ligament (bivalve)</span> Part of the bivalve shell

A hinge ligament is a crucial part of the anatomical structure of a bivalve shell, i.e. the shell of a bivalve mollusk. The shell of a bivalve has two valves and these are joined by the ligament at the dorsal edge of the shell. The ligament is made of a strong, flexible and elastic, fibrous, proteinaceous material which is usually pale brown, dark brown or black in color.

<span class="mw-page-title-main">Adductor muscles (bivalve)</span> Main muscular system in bivalves

The adductor muscles are the main muscular system in bivalve mollusks. In many parts of the world, when people eat scallops, the adductor muscles are the only part of the animal which is eaten. Adductor muscles leave noticeable scars or marks on the interior of the shell's valves. Those marks are often used by scientists who are in the process of identifying empty shells to determine their correct taxonomic placement.

<span class="mw-page-title-main">Ostreoidea</span> Superfamily of bivalves

Ostreoidea is a taxonomic superfamily of bivalve marine mollusc, sometimes simply identified as oysters, containing two families. The ostreoids are characterized in part by the presence of a well developed axial rod. Anal flaps are known to exist within the family Ostreidae but not within the more-primitive Gryphaeidae. The scar from the adductor muscle is simple, with a single, central scar. In the majority, the right valve is less convex than the left.

References

  1. Charles Doolittle Walcott (1886). Second contribution to the studies on the Cambrian faunas of North America. Vol. 30 of Geological Survey bulletin. Govt. Print. Off. pp. 1–369.
  2. 1 2 3 Carter, J.G.; et al. (2011). "A Synoptical Classification of the Bivalvia (Mollusca)" (PDF). Paleontological Contributions. 4: 1–47.
  3. 1 2 3 4 5 6 7 8 9 10 Elicki, O.; Gürsu, S. (2009). "First record of Pojetaia runnegari Jell, 1980 and Fordilla Barrande, 1881 from the Middle East (Taurus Mountains, Turkey) and critical review of Cambrian bivalves" (PDF). Paläontologische Zeitschrift. 83 (2): 267–291. doi:10.1007/s12542-009-0021-9. S2CID   49380913. Archived from the original (PDF) on 2013-10-29. Retrieved 2012-01-07.
  4. The Paleobiology Database Fordillidae entry accessed 4 January 2012
  5. 1 2 The Paleobiology Database Fordilla entry accessed 4 January 2012
  6. Pojeta, J. (1975). "Fordilla troyensis Barrande and early pelecypod phylogeny". Bulletins of American Paleontology. 67: 363–384. Archived from the original on 2012-01-06.
  7. 1 2 Vendrasco, M.J.; Checa, A.G.; Kouchinsky, A.V. (2011). "Shell microstructure of the early bivalve Pojetaia and the independent origin of nacre within the Mollusca". Palaeontology. 54 (4): 825–850. doi: 10.1111/j.1475-4983.2011.01056.x .
  8. Bouchet, Philippe; Rocroi, Jean-Pierre; Bieler, Rüdiger; Carter, Joseph G.; Coan, Eugene V. (2010). "Nomenclator of Bivalve Families with a Classification of Bivalve Families". Malacologia . 52 (2): 1–184. doi:10.4002/040.052.0201. S2CID   86546840.