Fordillidae

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Fordillidae
Temporal range: Tommotian–early Middle Cambrian
Fordilla troyensis shell.jpg
Fordilla troyensis outer shell surface [1]
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Mollusca
Class: Bivalvia
Order: Fordillida
Superfamily: Fordilloidea
Family: Fordillidae
Pojeta, 1975
Genera

Fordillidae is an extinct family of early bivalves [2] and one of two families in the extinct superfamily Fordilloidea. The family is known from fossils of early to middle Cambrian age found in North America, Greenland, Europe, the Middle East, Asia, and Australia. [3] [4] [5] The family currently contains two genera, Fordilla and Pojetaia , each with up to three described species. [3] [5] Due to the size and age of the fossil specimens, Fordillidae species are included as part of the Turkish Small shelly fauna. [3]

Description

Fordilla are small bivalves with valves that are equal in size and suboval in shape. In size Fordilla specimens reach a total shell length of up to 4 millimetres (0.16 in) and a height of 25 millimetres (0.98 in). [3] [6] The shells are compressed laterally and the back edge is slightly broadened. The rear adductor is less developed and smaller than the front adductor, while the small pedal retractor muscle scar is positioned near the front adductor scar. [3] The valve hinge is usually straight to slightly convexly curved and each valve will have at most one tooth present. The external surface of the shell occasionally show faint ribbing. [3]

Similar to Fordilla, species of Pojetaia are small, with valves to less than 2 millimetres (0.079 in) in length. Pojetaia species have an overall shape which is suboval, with the subequal valves slightly elongated. The ligament is straight with an umbo which is central to subcentral. In contrast to Fordilla the rear adductor muscle was larger and more developed then the front adductor, with pallial muscles arranged along the valve margins. Also in contrast to Fordilla, valves of Pojetaia possess between one and three teeth, with up to two teeth per valve. The exteriors of the shells show faint ribbing and fine comarginal growth lines. [3]

The inner shell layers of Fordilla and Pojetaia species both consist of layers of carbonate, which is akin to the laminar aragonite layer found in extant monoplacophora. [7] The structuring is similar to shell layering found in the extinct genera Anabarella and Watsonella which is thought to suggest that members of the phylum Mollusca developed nacre independently several times. [7]

Related Research Articles

<span class="mw-page-title-main">Bivalvia</span> Class of molluscs

Bivalvia, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bivalves have no head and they lack some usual molluscan organs, like the radula and the odontophore. The class includes the clams, oysters, cockles, mussels, scallops, and numerous other families that live in saltwater, as well as a number of families that live in freshwater. The majority are filter feeders. The gills have evolved into ctenidia, specialised organs for feeding and breathing. Most bivalves bury themselves in sediment, where they are relatively safe from predation. Others lie on the sea floor or attach themselves to rocks or other hard surfaces. Some bivalves, such as the scallops and file shells, can swim. Shipworms bore into wood, clay, or stone and live inside these substances.

<span class="mw-page-title-main">Craniata (brachiopod)</span> Class of marine lamp shells

Craniata is a class of brachiopods originating in the Cambrian period and still extant today. It is the only class within the subphylum Craniiformea, one of three major subphyla of brachiopods alongside linguliforms and rhynchonelliforms. Craniata is divided into three orders: the extinct Craniopsida and Trimerellida, and the living Craniida, which provides most information on their biology. Living members of the class have shells which are composed of calcite, though some extinct forms my have aragonite shells. The shells are inarticulate and are usually rounded in outline. There is no pedicle; the rear edge of the body cavity is a smooth and flat wall perforated by the anus. This class of brachiopods has an unsupported lophophore with only a single row of tentacles. In the absence of a pedicle, the shell is usually attached directly to a hard substrate. Many craniiforms are encrusting animals which attach directly to the shell of another animal, usually another brachiopod. The plicae from the host brachiopod will then appear within the shell of the craniiform.

<span class="mw-page-title-main">Bivalve shell</span> Seashell

A bivalve shell is part of the body, the exoskeleton or shell, of a bivalve mollusk. In life, the shell of this class of mollusks is composed of two hinged parts or valves. Bivalves are very common in essentially all aquatic locales, including saltwater, brackish water, and freshwater. The shells of bivalves commonly wash up on beaches and along the edges of lakes, rivers, and streams. Bivalves by definition possess two shells or valves, a "right valve" and a "left valve", that are joined by a ligament. The two valves usually articulate with one another using structures known as "teeth" which are situated along the hinge line. In many bivalve shells, the two valves are symmetrical along the hinge line—when truly symmetrical, such an animal is said to be equivalved; if the valves vary from each other in size or shape, inequivalved. If symmetrical front-to-back, the valves are said to be equilateral, and are otherwise considered inequilateral.

<span class="mw-page-title-main">Brachiopod</span> Phylum of marine animals also known as lamp shells

Brachiopods, phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachiopods, a stalk-like pedicle projects from an opening near the hinge of one of the valves, known as the pedicle or ventral valve. The pedicle, when present, keeps the animal anchored to the seabed but clear of sediment which would obstruct the opening.

Anabarella is a species of bilaterally-flattened monoplacophoran mollusc, with a morphological similarity to the rostroconchs. Its shell preserves evidence of three mineralogical textures on its outer surface: it is polygonal near the crest of the shell, subsequently changing to both spiny and stepwise. Its internal microstructure is calcitic and semi-nacreous. Its name reflects its provenance from Anabar, Siberia. It has been interpreted as ancestral to the rostroconchs, and has been aligned to the Helcionellidae.

<i>Fordilla</i> Extinct genus of bivalves

Fordilla is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early Cambrian fossils found in North America, Greenland, Europe, the Middle East, and Asia. The genus currently contains three described species, Fordilla germanica, Fordilla sibirica, and the type species Fordilla troyensis.

<span class="mw-page-title-main">Rhynchonelliformea</span> Subphylum of brachiopods

Rhynchonelliformea is a major subphylum and clade of brachiopods. It is roughly equivalent to the former class Articulata, which was used previously in brachiopod taxonomy up until the 1990s. These so-called articulated brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations and separate sets of simple opening and closing muscles.

Pojetaia is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early to Middle Cambrian fossils found in North America, Greenland, Europe, North Africa, Asia, and Australia. The genus currently contains two accepted species, Pojetaia runnegari, the type species, and Pojetaia sarhroensis, though up to seven species have been proposed. The genera Buluniella, Jellia, and Oryzoconcha are all considered synonyms of Pojetaia.

<i>Tellimya ferruginosa</i> Species of bivalve

Tellimya ferruginosa is a species of small marine bivalve mollusc in the family Lasaeidae. It is found on the eastern side of the Atlantic Ocean.

<span class="mw-page-title-main">Bakevelliidae</span> Extinct family of bivalves

Bakevelliidae is an extinct family of prehistoric bivalves that lived from the Late Mississippian until the Middle Eocene. Bakevelliidae species are found worldwide, excluding Antarctica. Living a stationary life attached to substrate in marine and brackish environments, they formed shells of an aragonite composition with a low amount of magnesium calcite. The family was named by William King in 1850. At least one genus in the family, Hoernesia, has a notably twisted commissure join.

Camya is an extinct genus of early bivalve and is the only genus in the extinct family Camyidae. The genus is known solely from early Middle Cambrian fossils found in Europe. The genus currently contains a solitary accepted species, Camya asy.

<span class="mw-page-title-main">Fordilloidea</span> Extinct superfamily of bivalves

Fordilloidea is an extinct superfamily of early bivalves containing two described families, Fordillidae and Camyidae and the only superfamily in the order Fordillida. The superfamily is known from fossils of early to middle Cambrian age found in North America, Greenland, Europe, the Middle East, Asia, and Australia. Fordillidae currently contains two genera, Fordilla and Pojetaia each with up to three described species while Camyidae only contains a single genus Camya with one described species, Camya asy. Due to the size and age of the fossil specimens, Fordillidae species are included as part of the Turkish Small shelly fauna.

Tuarangia is a Cambrian shelly fossil interpreted as an early bivalve, though alternative classifications have been proposed and its systematic position remains controversial. It is the only genus in the extinct family Tuarangiidae and order Tuarangiida. The genus is known solely from Middle to Late Cambrian fossils found in Europe and New Zealand. The genus currently contains two accepted species, Tuarangia gravgaerdensis and the type species Tuarangia paparua.

Concavodonta is an extinct genus of early bivalve in the extinct family Praenuculidae. The genus is one of three genera in the subfamily Concavodontinae. Concavodonta is known solely from late Ordovician, Caradoc epoch, fossils found in Europe and South America. The genus currently contains three accepted species, Concavodonta imbricata, Concavodonta ovalis and the type species Concavodonta ponderata.

Hemiconcavodonta is an extinct genus of bivalve in the extinct family Praenuculidae. The genus is one of three genera in the subfamily Concavodontinae. Hemiconcavodonta is known solely from late Ordovician, Caradoc epoch, fossils found in South America. The genus currently contains a single accepted species, Hemiconcavodonta minuta.

Praenuculinae is an extinct subfamily of prehistoric bivalves in the family Praenuculidae. Praenuculinae species lived from the middle Ordovician through the late Devonian. Praenuculinae fossils are found in Europe, Africa, North America and South America, and species are thought to have been stationary attached to substrate in shallow infaunal marine water environments where they formed shells of an aragonite composition. The subfamily Praenuculinae was named by Teresa M. Sánchez in 1999.

<i>Cucullaea labiata</i> Species of bivalve

Cucullaea labiata is a species of saltwater clam or ark shell, a marine bivalve mollusk in the family Cucullaeidae.

<span class="mw-page-title-main">Ligament (bivalve)</span>

A hinge ligament is a crucial part of the anatomical structure of a bivalve shell, i.e. the shell of a bivalve mollusk. The shell of a bivalve has two valves and these are joined by the ligament at the dorsal edge of the shell. The ligament is made of a strong, flexible and elastic, fibrous, proteinaceous material which is usually pale brown, dark brown or black in color.

<span class="mw-page-title-main">Adductor muscles (bivalve)</span> Main muscular system in bivalves

The adductor muscles are the main muscular system in bivalve mollusks. In many parts of the world, when people eat scallops, the adductor muscles are the only part of the animal which is eaten. Adductor muscles leave noticeable scars or marks on the interior of the shell's valves. Those marks are often used by scientists who are in the process of identifying empty shells to determine their correct taxonomic placement.

<span class="mw-page-title-main">Paterinata</span> Extinct class of marine lamp shells

Paterinata is an extinct class of linguliform brachiopods which lived from the lower Cambrian ("Tommotian") to the Upper Ordovician (Hirnantian). It contains the single order Paterinida and the subfamily Paterinoidea. Despite being some of the earliest brachiopods to appear in the fossil record, paterinides stayed as a relatively subdued and low-diversity group even as other brachiopods diversified later in the Cambrian and Ordovician. Paterinides are notable for their high degree of convergent evolution with rhynchonelliform (articulate) brachiopods, which have a similar set of muscles and hinge-adjacent structures.

References

  1. Charles Doolittle Walcott (1886). Second contribution to the studies on the Cambrian faunas of North America. Vol. 30 of Geological Survey bulletin. Govt. Print. Off. pp. 1–369.
  2. Carter, J.G.; et al. (2011). "A Synoptical Classification of the Bivalvia (Mollusca)" (PDF). Paleontological Contributions. 4: 1–47.
  3. 1 2 3 4 5 6 7 Elicki, O.; Gürsu, S. (2009). "First record of Pojetaia runnegari Jell, 1980 and Fordilla Barrande, 1881, from the Middle East (Taurus Mountains, Turkey) and critical review of Cambrian bivalves" (PDF). Paläontologische Zeitschrift. 83 (2): 267–291. doi:10.1007/s12542-009-0021-9. S2CID   49380913. Archived from the original (PDF) on 2013-10-29. Retrieved 2012-01-06.
  4. The Paleobiology Database Fordillidae entry accessed 4 January 2012
  5. 1 2 The Paleobiology Database Fordilla entry [ permanent dead link ] accessed 4 January 2012
  6. Pojeta, J. (1975). "Fordilla troyensis Barrande and early pelecypod phylogeny". Bulletins of American Paleontology. 67: 363–384. Archived from the original on 2012-01-06.
  7. 1 2 Vendrasco, M.J.; Checa, A.G.; Kouchinsky, A.V. (2011). "Shell microstructure of the early bivalve Pojetaia and the independent origin of nacre within the Mollusca". Palaeontology. 54 (4): 825–850. doi: 10.1111/j.1475-4983.2011.01056.x .