Girvanella Temporal range: | |
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Scientific classification ![]() | |
Domain: | Bacteria |
Phylum: | Cyanobacteria |
Class: | Cyanophyceae |
Order: | Oscillatoriales |
Genus: | † Girvanella Nicholson & Etheridge, 1878 |
Type species | |
Girvanella problematica Nicholson & Etheridge, 1878 emend. Wood, 1957 | |
Species | |
| |
Synonyms | |
NicholsoniaKorde, 1973 |
Girvanella is a fossil thought to represent the calcified sheath of a filamentous cyanobacterium known from the Burgess Shale [1] and other Cambrian fossil deposits. [2] Specimens are also known from the Early Ordovician San Juan Formation, Argentina. [3]
Girvanella was originally described as a foraminifera. [4] It was later assigned to the now-obsolete family porostromata. [5] In 2020, it was assigned to the order Oscillatoriales. [6]
Girvanella is characterised by having flexing, tubular filaments with a uniform diameter usually between 10 and 30 microns (rarely up to 100 microns). The walls of these tubules are relatively thick and calcareous. These tubules are typically (but not always) twisted together into nodules, and often encrust other objects including foraminifera. [4]
Fossils of Girvanella are found from the Cambrian through the Cretaceous. [4]
Girvanella fossils are found in a wide range of environmental conditions, most commonly shallow-shelf carbonate facies, but also in nonmarine limestones. Recent caliche deposits in Barbados may be referable to Girvanella. [4]
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At 508 million years old, it is one of the earliest fossil beds containing soft-part imprints.
Stromatolites or stromatoliths are layered sedimentary formations (microbialite) that are created mainly by photosynthetic microorganisms such as cyanobacteria, sulfate-reducing bacteria, and Pseudomonadota. These microorganisms produce adhesive compounds that cement sand and other rocky materials to form mineral "microbial mats". In turn, these mats build up layer by layer, growing gradually over time.
Ottoia is a stem-group archaeopriapulid worm known from Cambrian fossils. Although priapulid-like worms from various Cambrian deposits are often referred to Ottoia on spurious grounds, the only clear Ottoia macrofossils come from the Burgess Shale of British Columbia, which was deposited 508 million years ago. Microfossils extend the record of Ottoia throughout the Western Canada Sedimentary Basin, from the mid- to late- Cambrian. A few fossil finds are also known from China.
A microfossil is a fossil that is generally between 0.001 mm and 1 mm in size, the visual study of which requires the use of light or electron microscopy. A fossil which can be studied with the naked eye or low-powered magnification, such as a hand lens, is referred to as a macrofossil.
Receptaculites is the name-bearing genus for an extinct group of conspicuous benthic marine genera, the receptaculitids, that lived from the Early Ordovician through the Permian period, peaking in the Middle Ordovician. The group's phylogenetic origin has long been obscure, with some arguing that they were calcareous algae, probably of the order Dasycladales. Receptaculitids lived in warm, shallow seas, but consensus disagreeing. They have been described from all continents except Antarctica. In some areas they were important reef-formers, and they also occur as isolated specimens.
Oncolites are sedimentary structures composed of oncoids, which are layered structures formed by the growth of one or more species of microorganisms, usually containing cyanobacteria. Oncolites are very similar to stromatolites, but, instead of forming columns, they form approximately spherical structures. The oncoids often form around a central nucleus, such as a shell fragment, and a calcium carbonate structure is deposited by encrusting microbes. Oncolites are indicators of warm waters in the photic zone, but are also known in contemporary freshwater environments. These structures rarely exceed 10 cm in diameter.
Marrellomorpha are an extinct group of arthropods known from the Cambrian to the Early Devonian. They lacked mineralised hard parts, so are only known from areas of exceptional preservation, limiting their fossil distribution. The best known member is Marrella, with thousands of specimens found in the Cambrian aged Burgess Shale of Canada. The group is divided up into two major orders, Marrellida and Acercostraca. Marrellida is recognised by the possession of head shields with two or three pairs of elongate spine-like projections, and three pairs of uniramous appendages on the cephalon, while Acercostraca generally have large ovoid carapaces that cover the entire upper half of the body, and five pairs of uniramous cephalic appendages. Both groups have unbranched antennules and a segmented trunk with biramous appendages. Recent research has suggested the previously enigmatic Cambrian trilobite-like arthropods Skania and Primicaris belong to this group. Their phylogenetic position is uncertain, various studies have alternatively placed them in the Arachnomorpha as relatives of Artiopoda, as related to Mandibulata, or as stem group euarthropods. Some authors have proposed that they may be closely related to sea spiders (Pycnogonida) within Chelicerata though the cladistical support for such a relationship is relatively weak.
Eiffelia is an extinct genus of sponges known from the Middle Cambrian Burgess Shale as well as several Early Cambrian small shelly fossil deposits. It is named after Eiffel Peak, which was itself named after the Eiffel Tower. It was first described in 1920 by Charles Doolittle Walcott. It belongs in the Hexactinellid stem group. 60 specimens of Eiffelia are known from the Greater Phyllopod bed, where they comprise 0.11% of the community.
The extinct genus Hedstroemia was once thought to be a rivulariacean cyanobacterium. It forms oval blobs composed of sinuous tubes that have rounded outlines and occurs in conjunction with Solenopora.
Rothpletzella is a genus of calcimicrobe known from the Silurian of Gotland, the Devonian of France, as well as the Ordovician of China. It has been hypothesised to be a cyanobacterium, and shares morphological similarities with extant cyanobacteria. The genus is named in honor of August Rothpletz.
Wetheredella is a genus of calcimicrobes initially described from the Silurian of England, and subsequently reported from the Upper Ordovician to the end of the Carboniferous periods; its reefs are stated as being characteristic of the Ordovician-Silurian periods. Its taxonomic position is uncertain; it has been suggested to be a foraminiferan, a cyanobacterium or simply treated as a microproblematicum; Vachard & Cózar (2010) refer it to the Algospongia, a similarly controversial group that they assigned to the Protista but later, per Vachard, 2021, to Algae incertae sedis, in its own family and suborder (Wetheredellina) in the order Moravamminida. The genus is named in honor of the geologist Edward Wethered.
The Cathedral Formation is a stratigraphic unit in the southern Canadian Rockies of Alberta and British Columbia, on the western edge of the Western Canada Sedimentary Basin. It is a thick sequence of carbonate rocks of Middle Cambrian age. It was named for Cathedral Mountain in Yoho National Park by Charles Doolittle Walcott, the discoverer of the Burgess shale fossils.
The Laborcita Formation is a geologic formation in the Sacramento Mountains of New Mexico. It preserves fossils dating back to the late Pennsylvanian to early Permian.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2016.
Sphaerocodium is a fossil that represents the remains of bacteria in the phylum Cyanobacteria, often called blue-green algae.
Proaulopora is a Cambrian–Ordovician fossil genus of calcareous algae. It has been variously thought to belong to the green algae, red algae or cyanobacteria. It was originally established by the Russian paleontologist Aleksandr Grigoryevich Vologdin in 1937, for species known from the Lower Cambrian of the western Altai Mountains.
The Eldon Formation is a stratigraphic unit that is present on the western edge of the Western Canada Sedimentary Basin in the southern Canadian Rockies of southwestern Alberta and southeastern British Columbia. It is a thick sequence of massive, cliff-forming limestones and dolomites that was named for Eldon Switch on the Canadian Pacific Railway near Castle Mountain in Banff National Park by Charles Doolittle Walcott, who discovered the Burgess Shale fossils. The Eldon Formation was deposited during Middle Cambrian time, and it includes fossil stromatolites. The Eldon forms the scenic cliffs at the top of Castle Mountain, and can also be seen at Mount Yamnuska and other mountains in Banff and Yoho National Parks.
"Porostromata" is an antiquated form taxon that refers to fossil cyanobacteria. The term porostromate is also used as a descriptor of textures and microstructure of stromatolites and oncolites which contain tubules or other cellular structures.
"Spongiostromata" is an antiquated form taxon that refers primarily to fossil cyanobacteria. "Spongiostromate" is also used to describe stromatolites and oncolites that do not preserve clear tubules or other cellular microstructure.
Beresellaceae is an extinct family of organisms of uncertain affinity, sometimes placed within the Metazoa, but tentatively assigned to the green alga order Dasycladales. Beresellids were cosmopolitan and their fossils are found in strata ranging in age from the late Devonian to the early Permian.
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