Hemibos Temporal range: Late Pliocene - Middle Pleistocene | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Artiodactyla |
Family: | Bovidae |
Subfamily: | Bovinae |
Subtribe: | Bubalina |
Genus: | † Hemibos Falconer, 1865 |
Type species | |
†Hemibos triquetricornis Falconer, 1865 | |
Species [1] | |
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Hemibos is an extinct even-toed ungulate, belonging to the family Bovidae. Its fossil remains were found across Asia and Europe, including findings from China, the Italian Peninsula, the Iberian Peninsula, the State of Palestine, Israel, Pakistan, and India. [2] [3] [4] [5]
This animal was a large bovid similar to the modern Asian buffalo (Bubalus bubalis), of which it is probably ancestral. It was characterized by elongated and non-prominent frontal bones; the bone cores of the horns did not have a neck at the base, and the angle between the horns was variable but usually between 85 ° and 110 °. Also, the orientation of the horns was variable according to the species: the species H. triquetricornis, H. acuticornis and H. galerianus possessed horns directed backwards, outwards and slightly upwards, while H. antelopinus and H. palaestinus possessed different morphologies. [6] The teeth were hypsodont, with increasing development of dental cement; the upper molars were square.
The genus Hemibos was first described in 1865 by Rütimeyer, based on fossils found in India in Plio-Pleistocene soils. The genus includes five species; three of these (H. acuticornis, H. triquetricornis and H. antelopinus) come from the Pinjor formation of the Siwaliks (Plio-Pleistocene, Indian subcontinent), [2] one (H. gracilis) is known from Gansu (China) in the lower Pleistocene, and another (H. galerianus) is the largest and most recent and comes from Ponte Galeria and Ponte Milvio (Rome, Italy) and dates from the Early Pleistocene/Middle Pleistocene boundary. In lower Pleistocene Spain, a form similar to H. gracilis was found. Another species attributed to Hemibos is H. palaestinicus from Israel, but the dating is not certain and a morphological analysis indicates it may belong to the genus Bison . The species Probubalus occipitalis is considered identical to H. triquetricornis.
Bos is a genus of bovines, which includes, among others, wild and domestic cattle.
Bubalus is a genus of Asiatic bovines that was proposed by Charles Hamilton Smith in 1827. Bubalus and Syncerus form the subtribe Bubalina, the true buffaloes.
Boselaphus is a genus of bovid. The nilgai is the sole living representative, although one other species is known from the fossil record.
Pachycrocuta is an extinct genus of prehistoric hyenas. The largest and most well-researched species is Pachycrocuta brevirostris, colloquially known as the giant short-faced hyena as it stood about 90–100 cm (35–39 in) at the shoulder and it is estimated to have averaged 110 kg (240 lb) in weight, approaching the size of a lioness, making it the largest known hyena. Pachycrocuta first appeared during the late Miocene. By 800,000 years ago, it became locally extinct in Europe, with it surviving in East Asia until at least 500,000 years ago, and possibly later elsewhere in Asia.
Pelorovis is an extinct genus of African wild cattle which existed during the Pleistocene epoch. The best known species is Pelorovis oldowayensis from Olduvai Gorge in Tanzania, from the Early Pleistocene. The species "Pelorovis" antiquus from the Late Pleistocene-Holocene has since been moved into Syncerus, the same genus as living African buffalo.
Bos acutifrons is the most ancient representative of the genus Bos cattle. Fossils of an individual of B. acutifrons were found in middle Pleistocene-aged strata of Siwalik Hills of Kashmir, in either modern Pakistan or India, in the 19th century. The prehistoric species was described, along with Bos planifrons, by Richard Lydekker in 1877. In 1898 Lydekker synonymised B. planifrons with B. acutifrons, reconsidering the skull found to be that of a female individual of the same species.
The tribe Bovini or wild cattle are medium to massive bovines that are native to Eurasia, North America, and Africa. These include the enigmatic, antelope-like saola, the African and Asiatic buffalos, and a clade that consists of bison and the wild cattle of the genus Bos. Not only are they the largest members of the subfamily Bovinae, they are the largest species of their family Bovidae. The largest species is the gaur, weighing up to 1,500 kg (3,300 lb).
Xenocyon is an extinct group of canids, either considered a distinct genus or a subgenus of Canis. The group includes Canis (Xenocyon) africanus, Canis (Xenocyon) antonii and Canis (Xenocyon) falconeri that gave rise to Canis (Xenocyon) lycanoides. The hypercarnivorous Xenocyon is thought to be closely related and possibly ancestral to modern dhole and the African wild dog, as well as the insular Sardinian dhole.
Lycaon is a genus of canid which includes the African wild dog and the extinct species Lycaon sekowei and Lycaon magnus.
Soergelia is a genus of extinct ovibovine caprine that was common across Europe, North America and Asia in the Pleistocene epoch.
Praemegaceros is an extinct genus of deer, known from the Pleistocene and Holocene of Western Eurasia. Praemegaceros is considered to be a genus of "giant deer", with many species having an estimated body mass of around 400 kilograms (880 lb), considerably larger than most living deer.
Hippopotamus behemoth is an extinct species of hippopotamus from the Early Pleistocene of the Levant. Fossils of it, and its probable ancestor, H. gorgops, are found in the ‘Ubeidiya site in the southern Levant, dating to the Early Pleistocene, around 1.4 million years ago.
Bubalina is a subtribe of wild cattle that includes the various species of true buffalo. Species include the African buffalo, the anoas, and the wild water buffalo. Buffaloes can be found naturally in sub-Saharan Africa, South Asia and Southeast Asia, and domestic and feral populations have been introduced to Europe, the Americas, and Australia. In addition to the living species, bubalinans have an extensive fossil record where remains have been found in much of Afro-Eurasia.
Bovina is a subtribe of the Bovini tribe that generally includes the two living genera, Bison and Bos. However, this dichotomy has been challenged recently by molecular work that suggests that Bison should be regarded as a subgenus of Bos. Wild bovinans can be found naturally in North America and Eurasia.
Capra dalii is a fossil species of goat discovered in Georgia in 2006. It is named for the Georgian goddess Dali, who was considered the guardian of hoofed animals such as ibexes and goats. Fragments of C. dalii fossils were first located at the Dmanisi archaeological site, and are believed to be related to the west Caucasian tur, Capra caucasica. The species is believed to have existed during the Early Pleistocene, around 1.76 million years ago, making it the oldest known example of the Capra genus.
Sivapardus is an extinct, little-known genus of feline with only one species assigned to it, Sivapardus punjabiensis. It was described in 1969 by the paleontologist Abu Bakr based on a partial mandible from the Upper Siwaliks in Pakistan; the locality it was found at is estimated to be from the Late Pliocene to Early Pleistocene. S. punjabiensis was a large cat with a short and broad snout that may have lived on open grasslands.
Sivacobus is an extinct species of antelope that lived in South Asia during the Plio-Pleistocene.
Boselaphus namadicus is an extinct species of bovid that lived in South Asia from the Late Pliocene to the Mid Pleistocene.
Megalovis is an extinct genus of bovid that lived in Eurasia during the Plio-Pleistocene.
Bos buiaensis is an extinct species of cattle. The species is known from a million year old skull fossil found at the archaeological site of Buya, Eritrea in 2003. It was reassembled by excavators from over one hundred shards.