Hollardops

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Hollardops
Temporal range: Emsian–Eifelian
Hollardops mesocristata, Early Devonian, TazoulaOt Formation, Jbel OufatEne & Jbel Issoumour, MaOder Region, Morocco - Houston Museum of Natural Science - DSC01602.JPG
Hollardops sp.
Houston Museum of Natural Science - DSC01602
Scientific classification
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Hollardops

Morzadec, 1997
Species
  • Asteropyge mesocristataLe Maître, 1952
  • H. aithassainorumChatterton et al., 2006
  • H. angustifronsVan Viersen & Kloc, 2022
  • Rhenops babiniMorzadec, 1983
  • Philipsmithiana burtandmimiaeLieberman & Kloc, 1997
  • H. boudibensisMorzadec, 2001
  • Rhenops circumapodemusSmeenk, 1983
  • Philipsmithiana hyfinkeliLieberman & Kloc, 1997
  • H. kyriarchosVan Viersen & Kloc, 2022
  • H. klugiVan Viersen & Kloc, 2022
  • H. luscusVan Viersen & Kloc, 2022
  • H. multatuliVan Viersen & Kloc, 2022
  • Greenops struveiMorzadec, 1969

Hollardops is a genus of trilobite in the order Phacopida that lived during the Devonian. Their fossils are found in the upper Emsian of western Europe (France, Spain) and in the lower Emsian to lowermost Eifelian of North Africa (Morocco, Algeria). The type species, Asteropyge mesocristata, was described from Algeria by Le Maître in 1952. The genus Hollardops was erected by Morzadec in 1997. [1] In the same year, Lieberman & Kloc [2] erected Modellops and Philipsmithiana but those genera are regarded as subjective synonyms of Hollardops. The 10-segmented thoracic condition of Hollardops is a rare feature among acastid trilobites that almost always have 11 thorax segments. Van Viersen & Kloc (2022) [3] revisited Hollardops and described a number of new species from the Devonian of Morocco. They also regarded Pennarbedops Bignon & Crônier, 2013 as a synonym of Hollardops. Van Viersen & Kloc construed Hollardops as a scavenger or predator with well-developed eyes, that used its shovel-like cephalon to plough the top layers of the sediment in search of food. Hollardops had small pits horizontally along the fringe of the exoskeleton; these are believed to have housed setae that allowed the trilobite to closely monitor its surroundings.

Description

Hollardops sp.
Houston Museum of Nat. Sci. - DSC01611
Both specimen are from Tazoulait Formation at
Jbel (Jebel) Oufatene
.mw-parser-output .geo-default,.mw-parser-output .geo-dms,.mw-parser-output .geo-dec{display:inline}.mw-parser-output .geo-nondefault,.mw-parser-output .geo-multi-punct,.mw-parser-output .geo-inline-hidden{display:none}.mw-parser-output .longitude,.mw-parser-output .latitude{white-space:nowrap}
30deg50'15''N 4deg54'07''W / 30.8374368degN 4.9018067degW / 30.8374368; -4.9018067 (Jbel Oufatene)
and Issimour
30deg58'01''N 5deg02'14''W / 30.9669834degN 5.0373266degW / 30.9669834; -5.0373266 (Jbel Issimour)
SE of Alnif, western of Oued Alnif, Ma'ider region, Morocco Hollardops mesocristata, Early Devonian, TazoulaOt Formation, Jbl OutfatEne & Jbel Issoumour, MaOder Region, Morocco - Houston Museum of Natural Science - DSC01611.JPG
Hollardops sp.
Houston Museum of Nat. Sci. - DSC01611
Both specimen are from Tazoulait Formation at
Jbel (Jebel) Oufatène 30°50′15″N4°54′07″W / 30.8374368°N 4.9018067°W / 30.8374368; -4.9018067 (Jbel Oufatène)
and Issimour 30°58′01″N5°02′14″W / 30.9669834°N 5.0373266°W / 30.9669834; -5.0373266 (Jbel Issimour)
SE of Alnif, western of Oued Alnif, Ma'ider region, Morocco

Hollardops have schizochroal eyes and a glabella that is slightly raised on the surface of the cephalon. Genal spines extend from the cephalon and extend to approximately the 6th thoracic segment.

Hollardops has 10 thoracic segments and also has 5 pairs of pleural lappets or spines projecting from the pygidium.

Length ranges from approximately 3 to 9 cm.

Related Research Articles

<span class="mw-page-title-main">Trilobite</span> Class of extinct, Paleozoic arthropods

Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.

<span class="mw-page-title-main">Harpetida</span> Extinct order of trilobites

Harpetida is one of the eleven orders of the extinct arthropod class Trilobita. The first harpetid trilobites appear in the Upper Cambrian, and the last species die out at the end of the Devonian period.

<span class="mw-page-title-main">Corynexochida</span> Extinct order of trilobites

Corynexochida is an order of trilobite that lived from the Lower Cambrian to the Late Devonian. Like many of the other trilobite orders, Corynexochida contains many species with widespread characteristics.

<i>Phacops</i> Genus of arthropods (fossil)

Phacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived in Europe, northwestern Africa, North and South America and China from the Late Ordovician until the very end of the Devonian, with a broader time range described from the Late Ordovician. It was a rounded animal, with a globose head and large eyes, and probably fed on detritus. Phacops is often found rolled up ("volvation"), a biological defense mechanism that is widespread among smaller trilobites but further perfected in this genus.

<i>Dalmanites</i> Extinct genus of trilobites

Dalmanites is a genus of trilobite in the order Phacopida. They lived from the Late Ordovician to Middle Devonian.

<i>Greenops</i> Extinct genus of trilobite

Greenops is a mid-sized Devonian trilobite of the order Phacopida, subfamily Asteropyginae. They are mainly reported from the mid-Devonian Hamilton Group of upstate New York and southwestern Ontario. A similar-looking trilobite from Morocco is often mis-labelled Greenops. Greenops had schizocroidal eyes, large genal spines and short, sharp spines at the tip of each segment of the pygidium ("tail"). Greenops lived in warm, fairly deep water. In the Hamilton Group of New York, they are found with Eldredgeops, Dipleura and Bellacartwrightia, a trilobite that resembles Greenops but has much larger pygidial spines. In Ontario, they are found in the Widder Formation, which outcrops at Arkona, where they are, by far, the dominant trilobite.

<i>Walliserops</i>

Walliserops is a genus of spinose phacopid trilobite, of the family Acastidae, found in Lower to Middle Devonian age rocks from the Anti-Atlas Mountains of Morocco. All species of Walliserops possess a three-pronged "trident" that protrudes from the glabella. Walliserops is most closely related to the genus Comura.

<i>Acimetopus</i>

Acimetopus Rasetti, 1966, is a genus of Eodiscinid trilobite belonging to the family Weymouthiidae Kobayashi (1943), Order Agnostida Salter (1864). It lived during the Botomian stage. = late Lower Cambrian Stage 4 ; the upper Botomian boundary corresponds to base of the Middle Cambrian, Miaolingian Series and Wuliuan stage.

<i>Meteoraspis</i>

Meteoraspis is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae. The various species lived from 501 to 490 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Meteoraspis are characteristic of Late Cambrian strata in North America, though they are found in Late Cambrian strata elsewhere in the world, such as M. nevensis from Victoria Land, Antarctica.

<i>Erbenochile</i>

Erbenochile is a genus of spinose phacopid trilobite, of the family Acastidae, found in Lower to Middle Devonian age rocks from Algeria and Morocco. Originally described from an isolated pygidium, the first complete articulated specimen of E. erbeni revealed the presence of extraordinarily tall eyes:

"Straight-sided towers of lenses... with [up to] 18 lenses in a vertical file"

<i>Cyphaspis</i> Extinct genus of trilobites

Cyphaspis is a genus of small trilobite that lived from the Late Ordovician to the Late Devonian. Fossils have been found in marine strata in what is now Europe, Africa and North America. Various species had a compact body, and a large, bulbous glabellum. Many species had long spines arranged similarly to closely related genera, such as Otarian, Otarionella, Chamaeleoaspis, and Namuropyge.

<i>Morocconites</i>

Morocconites malladoides is an average size trilobite, which lived during the Devonian period, in what is now southern Morocco. This species is assumed to be a close relative of Acastoides. The most conspicuous feature is the very long upcurved frontal medial spine, a bit like an avocet bill. It is the only known species in this genus.

<span class="mw-page-title-main">Holmiidae</span> Family of trilobites

Holmiidae is a family of trilobites, that lived during the Lower Cambrian (Atdabanian). The Holmiidae is a diverse family of eight genera containing at least 17 species. It includes some of the earliest trilobites of Baltica. Holmiidae occur throughout Baltica and Western Laurentia, and also in Morocco.

<i>Odontochile</i> Genus of trilobites

Odontochile is a genus of trilobites in the order Phacopida, family Dalmanitidae.

<i>Symphysops</i> Extinct genus of trilobites

Symphysops is a genus of trilobites of average size, belonging to the Cyclopygidae family. It had a cosmopolitan distribution and lived from the Middle to the Upper Ordovician. It has been found in Canada, China, the Czech Republic (Bohemia), Iran, Ireland, Kazakhstan, Poland, Morocco, Spain, Scotland and Wales. The name Symphysops refers to the fused eyes, common to the species of this genus. Some (sub)species of the cyclopygid genera Cyclopyge and Pricyclopyge share this character, but Symphysops uniquely combines the merged eye with a frontal thorn on the head and the "lower eyelid".

<i>Coltraneia</i>

Coltraneia is a genus of trilobite, that lived during the upper Emsian and lower Eifelian, and has been found in Algeria, France, Germany, Morocco and Spain.

<i>Minicryphaeus</i>

Minicryphaeus is a genus of trilobite, that lived during the Pragian in what is today Morocco (Anti-Atlas).

<i>Psychopyge</i>

Psychopyge is a genus of trilobite, that lived during the upper Emsian and has been found in Germany and Morocco. It is characterized by the swordlike extension from the front of the head.

<i>Gerastos</i> Extinct genus of trilobites

Gerastos is a genus of proetid trilobite in the family Proetidae that lived between the Pragian and Eifelian of the Lower-Middle Devonian, spanning approximately 21 million years.

Tolkienia is an extinct genus of trilobite in the family Acastidae. It consists of three species split from Comura based on morphological differences. The genus is was present during the Emsian stage of the Early Devonian epoch around 400 Ma.

References

  1. Morzadec, Pierre (1997-04-29). "Les Trilobites Asteropyginae du Dévonien de l'Ougarta (Algérie)". Palaeontographica Abteilung A (in French): 143–158. doi:10.1127/pala/244/1997/143.
  2. Lieberman, Bruce S.; Kloc, Gerald J. (1997). "Evolutionary and biogeographic patterns in the Asteropyginae (Trilobita, Devonian) Delo, 1935. Bulletin of the AMNH ; no. 232".{{cite journal}}: Cite journal requires |journal= (help)
  3. Viersen, Allart P. VAN; J. Kloc, Gerald (2022-12-09). "Functional morphology, coaptation and palaeoecology of Hollardops (Trilobita, Acastidae), with descriptions of new species and two new genera from the Devonian of Morocco". Geologica Belgica. doi: 10.20341/gb.2022.005 . ISSN   1374-8505.
  4. Google Maps: Jebel Oufatene
  5. Google Maps: [Jebel] Issimour