Homatula

Homatula
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Cypriniformes
Family:	Nemacheilidae
Genus:	Homatula Nichols, 1925
Type species
Nemachilus potanini Günther, 1896 [1]

Homatula is a genus of stone loaches endemic to China.

Species

These are the currently recognized species in this genus: ^{[2] [3]}

• Homatula acuticephala (W. Zhou & J. C. He, 1993)
• Homatula anguillioides (S. Q. Zhu & S. H. Wang, 1985)
• Homatula anteridorsalis Li, Che & Zhou, 2019
• Homatula berezowskii (Günther, 1896)
• Homatula change Endruweit, 2015 ^[4]
• Homatula coccinocola Endruweit, Min & Yang, 2018
• Homatula cryptoclathrata Li, Che & Zhou, 2019
• Homatula disparizona R. Min, J. X. Yang & X. Y. Chen, 2013 ^[5]
• Homatula dotui Nguyen, Wu, Cao & Zhang, 2021
• Homatula erhaiensis (S. Q. Zhu & W. X. Cao, 1988)
• Homatula gelao Xiao, 2025
• Homatula geminusclathrata Li, Yang, Guo & Zhou, 2022
• Homatula guanheensis Zhou, Ma, Wang, Tang, Meng & Nie, 2021
• Homatula laxiclathra J. H. Gu & E. Zhang, 2012 ^[6]
• Homatula longibarbata Li, Yang, Guo & Zhou, 2022
• Homatula longidorsalis (J. X. Yang, Y. R. Chen & Kottelat, 1994)
• Homatula microcephala Li, Yang, Guo & Zhou, 2022
• Homatula nanpanjiangensis (R. Min, X. Y. Chen & J. X. Yang, 2010)
• Homatula nigra Li, Che & Zhou, 2019
• Homatula oligolepis (W. X. Cao & S. Q. Zhu, 1989)
• Homatula oxygnathus (Regan, 1908)
• Homatula potanini (Günther, 1896)
• Homatula pycnolepis Y. T. Hu & E. Zhang, 2010
• Homatula robusta Min, Zhao, Shi & Yang, 2022
• Homatula shexiang Cao, Liu, Zeng & Zhang, 2025
• Homatula submarginata Endruweit, 2025
• Homatula tigris Che, Dao, Chen, Pan, Hua, Liang & Wang, 2023
• Homatula variegata (Dabry de Thiersant, 1874)
• Homatula wenshanensis C. Li, H. Yang, W. Li & T. Liu, 2017
• Homatula wujiangensis R. H. Ding & Q. X. Deng, 1990
• Homatula wuliangensis R. Min, J. X. Yang & X. Y. Chen, 2012 ^[7]
• Homatula xiangzhi Cao, Liu, Zeng & Zhang, 2025

References

1. Fricke, Ron; Eschmeyer, William N. & van der Laan, Richard (eds.). "Genera in the family Nemacheilidae". Catalog of Fishes . California Academy of Sciences . Retrieved 4 December 2024.
2. Fricke, Ron; Eschmeyer, William N. & van der Laan, Richard (eds.). "Species in the genus Homatula". Catalog of Fishes . California Academy of Sciences . Retrieved 11 November 2025.
3. Kottelat, M. (2012). "Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei)" (PDF). The Raffles Bulletin of Zoology (Suppl. No. 26): 1–199. Archived from the original (PDF) on 2013-02-11.
4. Endruweit, M. (2015). "Homatula change, a new nemacheilid loach from the upper Black River basin in Yunnan, China (Teleostei: Nemacheilidae)". Ichthyological Exploration of Freshwaters. 26 (1): 65–72.
5. Min, R.; Yang, J.-X.; Chen, X.-Y. (2013). "Homatula disparizona, a new species of loach from the Red River drainage in China (Teleostei: Nemacheilidae)". Ichthyological Exploration of Freshwaters. 23 (4): 351–355.
6. Gu, J.-H.; Zhang, E. (2012). "Homatula laxiclathra (Teleostei: Balitoridae), a new species of nemacheiline loach from the Yellow River drainage in Shaanxi Province, northern China". Environmental Biology of Fishes. 94 (4): 591–599. Bibcode:2012EnvBF..94..591G. doi:10.1007/s10641-011-9965-1.
7. Min, R.; Yang, J.-X.; Chen, X.-Y. (2012). "Homatula wuliangensis (Teleostei: Nemacheilidae), a new loach from Yunnan, China" (PDF). Zootaxa. 3586: 313–318. doi:10.11646/zootaxa.3586.1.29.