The Chinle Formation is an Upper Triassic continental geologic formation in the western United States which has yielded fossils of many archosaurs: a group of vertebrates that includes crocodiles, pterosaurs, dinosaurs (including birds), and other extinct relatives. This is a list of all archosaurs from the formation, including suchians (crocodilian relatives) and avemetatarsalians (dinosaurs, pterosaurs, and their relatives), as well as the abundant crocodilian-like phytosaurs, which may either be true archosaurs or very close relatives of Archosauria.
Phytosaurs of the Chinle Formation | ||||||
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Genus | Species | State | Member | Abundance | Notes | Images |
L. crosbiensis [1] |
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S. adamanensis [1] |
| Known from a single skull | Previously referred to Leptosuchus, actually referable to Smilosuchus. [3] | |||
S. gregorii |
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| Known from a handful of skulls. | A basal leptosuchomorph. | ||
S. lithodendrorum [3] |
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| Known from only one skull. | Previously considered synonymous with S. adamanensis. | ||
cf. Paleorhinus [4] | Indeterminate [4] |
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M. buceros [5] |
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M. mccauleyi [5] |
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M. pristinus [5] |
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Pravusuchus [3] | P. hortus |
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| Three skulls. | A leptosuchomorph phytosaur closely related to Mystriosuchini [3] [6] or Nicrosaurus . [7] | |
Protome [8] | P. batalaria |
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| Partial skull and jaw. | A phytosaur originally described as a juvenile Leptosuchus , but probably an unrelated non-leptosuchomorph [8] or mystriosuchin. [7] | |
R. gregorii [9] | A large late-surviving mystriosuchin phytosaur closely related to Machaeroprosopus (formerly known as Pseudopalatus ), [10] and often considered a subgenus of that taxon. [2] [6] [7] | |||||
Color key
| Notes Uncertain or tentative taxa are in small text; |
Aetosaurs of the Chinle Formation | ||||||
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Genus | Species | State | Member | Abundance | Notes | Images |
Adamanasuchus | A. eisenhardtae |
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| Osteoderms and skull and postcranial fragments. | A basal desmatosuchine related to Calyptosuchus and Scutarx . [11] | |
C. wellesi [13] |
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| Common, although "non-armor material is comparatively rare". [13] | A basal desmatosuchine originally considered a species of Stagonolepis . | ||
D. spurensis [15] |
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| Numerous osteoderms and other skeletal material | A large desmatosuchin which was fairly common in the lower part of the Chinle Formation. Prior to 2008, fossils of D. spurensis were listed as the species D. haplocerus, until it was determined that D. haplocerus was a species name originally based on undiagnostic fossils. [11] | ||
D. smalli [16] |
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| A single lateral osteoderm. | A desmatosuchin which is very rare in the Chinle Formation. One of the youngest known occurrences of the genus Desmatosuchus. [11] | ||
P. andressorum [17] |
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| Osteoderms | A significant number of researchers feel that the North American material referred to the German Paratypothorax species P. andressorum by Hunt and Lucas in 1992 is actually distinct. [17] | ||
Indeterminate [18] |
| Osteoderms and minor postcranial fragments. | A paratypothoracin typothoracine known from isolated material similar to more complete specimens from Germany. [19] | |||
R. chamaensis |
| Osteoderms and possible ankle and vertebrae remains. | A late-surviving paratypothoracin typothoracine. Originally referred to Desmatosuchus, but later reinterpreted as a relative of Paratypothorax based on a phylogenetic analysis performed on the remains by Parker in 2003. [20] New genus names were erected independently by Parker (coining the name Heliocanthus) and Lucas et al. (coining the name Rioarribasuchus), prompting an investigation into reports of unethical taxonomic claim-jumping. | |||
Scutarx [11] | S. deltatylus [11] |
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| Four partial skeletons including skull fragments. | A large basal desmatosuchine based on fossils originally referred to "Stagonolepis" (=Calyptosuchus) wellesi, a close relative. Fossils of Scutarx represent the highest stratigraphic occurrence of Stagonolepis-like aetosaurs in Petrified Forest National Park. | |
Stenomyti | S. huangae |
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| Partial skeleton including a well-preserved skull. | A small aetosaurine similar to Aetosaurus . | |
T. coccinarum [21] |
| Common in the Owl Rock and Petrified Forest Members. [21] | A large, wide-bodied typothoracine abundant throughout the Chinle Formation. Almost all of the known Chinlean Typothorax fossils are isolated whole or fragmentary osteoderms. [21] | |||
Tecovasuchus [22] | T. chatterjeei |
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| Osteoderms. | Previously thought to be more widespread in the formation. [22] | |
Kryphioparma [22] | K. caerula |
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| Osteoderms. | Described in 2023, probably related to other typothoracines. [22] | |
Crocodylomorphs of the Chinle Formation | ||||||
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Genus | Species | State | Member | Abundance | Notes | Images |
H. agilis [23] |
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P. mccreai [24] |
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Paracrocodylomorphs of the Chinle Formation | ||||||
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Genus | Species | State | Member | Abundance | Notes | Images |
E. okeeffeae |
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| A six-foot-long shuvosaurid from the Coelophysis quarry of Ghost Ranch. | |||
P. gracilis [25] |
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P. kirkpatricki [26] |
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Indeterminate [27] |
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| Jaw fragment. | Identified from a fossil which has been reinterpreted as an indeterminate "rauisuchian". | ||
S. inexpectatus [28] |
| Following a rediagnosis of the genus and species, only shuvosaurid fossils from the Cooper Canyon Formation are reliably identifiable as Shuvosaurus itself. [29] | ||||
V. haydeni [30] |
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| A large rauisuchid similar to Postosuchus . | |||
Miscellaneous suchians of the Chinle Formation | ||||||
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Genus | Species | State | Member | Abundance | Notes | Images |
Acaenasuchus [31] | A. geoffreyi [31] |
| Known from a handful of isolated osteoderms. [31] | A controversial suchian based primarily on osteoderms. Some paleontologists have considered Acaenasuchus to be juvenile Desmatosuchus haplocerus scutes, while others considered it a separate genus of small aetosaurs. [31] Most recently it has been reconsidered an aetosauriform related to, but not within, Aetosauria. It is also related to Revueltosaurus (also from the Chinle) and the armored reptile Euscolosuchus from Virginia. [32] | ||
R. callenderi [34] |
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R. hunti [35] |
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Color key
| Notes Uncertain or tentative taxa are in small text; |
Basal Dinosauriforms of the Chinle Formation | ||||||
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Genus | Species | State | Member | Abundance | Notes | Images |
Dromomeron | D. romeri |
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| A lagerpetid pterosauromorph found at various Ghost Ranch quarries and elsewhere in the Chinle Formation. | ||
E. baldwini |
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| A silesaurid dinosauriform closely related to the Polish genus Silesaurus . [37] | |||
K. williamparkeri |
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| A silesaurid dinosauriform with adaptations for herbivory. | |||
Prosauropod tracks are present in the Redonda, Sloan Canyon, and Sheep Pen Sandstone formations. Possibly the Rock Point Formation as well. Geographically, the tracks are present in New Mexico. [36]
Theropod tracks have been found in Utah and New Mexico recovered from the Redonda, Sloan Canyon, and Sheep Pen Sandstone formations. [36] Indeterminate theropod remains are stratigraphically present in the Petrified Forest, Bluewater Creek, and Rock Point members of New Mexico. [36]
Theropods and near-relatives of the Chinle Formation | ||||||
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Genus | Species | State | Member | Abundance | Notes | Images |
C. arizonensis [36] |
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| A coelophysine coelophysid | |||
C. bryansmalli [36] |
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| A large saurischian alternatively considered a herrerasaurid or a theropod related to Tawa hallae . | |||
C. bauri |
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C. sp. [40] |
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| Originally assigned to C. bauri, but likely a different taxon. [40] | |||
C. longicollis [36] |
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| Now considered a junior synonym of C. bauri. | |||
C. willistoni |
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| Now considered a junior synonym of C. bauri. | |||
D. chauliodus |
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| Skull and neck vertebrae fragments | A possible basal theropod with a short skull and enlarged teeth. | ||
T. hallae |
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| A probable basal theropod showing a mosaic of features similar to neotheropods and herrerasaurids. | |||
Phytosaurs are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. and are sometimes referred to as parasuchians. Phytosauria, Parasuchia, Parasuchidae, and Phytosauridae have often been considered equivalent groupings containing the same species. Some recent studies have offered a more nuanced approach, defining Parasuchidae and Phytosauridae as nested clades within Phytosauria as a whole. Phytosaurs were long-snouted and heavily armoured, bearing a remarkable resemblance to modern crocodilians in size, appearance, and lifestyle, as an example of convergence or parallel evolution
Postosuchus, meaning "Crocodile from Post", is an extinct genus of rauisuchid reptiles comprising two species, P. kirkpatricki and P. alisonae, that lived in what is now North America during the Late Triassic. Postosuchus is a member of the clade Pseudosuchia, the lineage of archosaurs that includes modern crocodilians. Its name refers to Post Quarry, a place in Texas where many fossils of the type species, P. kirkpatricki, were found. It was one of the apex predators of its area during the Triassic, larger than the small dinosaur predators of its time. It was a hunter which probably preyed on large bulky herbivores like dicynodonts and many other creatures smaller than itself.
Eucoelophysis is a genus of silesaurid dinosauriform from the Late Triassic (Norian) period Chinle Formation of New Mexico. It was assumed to be a coelophysid upon description, but a study by Nesbitt et al. found that it was actually a close relative of Silesaurus, which was independently supported by Ezcurra (2006), who found it to be the sister group to Dinosauria, and Silesaurus as the next most basal taxon.
Aetosaurs are heavily armored reptiles belonging to the extinct order Aetosauria. They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs more closely related to crocodilians than to birds and other dinosaurs. All known aetosaurs are restricted to the Late Triassic, and in some strata from this time they are among the most abundant fossil vertebrates. They have small heads, upturned snouts, erect limbs, and a body ornamented with four rows of plate-like osteoderms. Aetosaur fossil remains are known from Europe, North and South America, parts of Africa, and India. Since their armoured plates are often preserved and are abundant in certain localities, aetosaurs serve as important Late Triassic tetrapod index fossils. Many aetosaurs had wide geographic ranges, but their stratigraphic ranges were relatively short. Therefore, the presence of particular aetosaurs can accurately date a site in which they are found.
Revueltosaurus is an extinct genus of suchian pseudosuchian from Late Triassic deposits of New Mexico, Arizona and North Carolina, United States. Many specimens, mostly teeth, have been assigned to Revueltosaurus over the years. Currently, three species are included in this genus, all of which were originally thought to represent monospecific genera of basal ornithischian dinosaurs. Revueltosaurus was about 1 meter long.
Spinosuchus is an extinct genus of trilophosaurid allokotosaur from the Late Triassic of Texas, southern United States. It has been assigned to a variety of groups over its history, from coelophysid dinosaur to pseudosuchian to uncertain theropod dinosaur and to Proterosuchidae. This uncertainty is not unusual, given that it was only known from a poorly preserved, wall-mounted, partial vertebral column of an animal that lived in a time of diverse, poorly known reptile groups. However, newly collected material and recent phylogenetic studies of early archosauromorphs suggest that it represents an advanced trilophosaurid very closely related to Trilophosaurus.
Krzyzanowskisaurus is the name given to a genus of archosaur from the Late Triassic-aged Chinle Formation and it is a tooth taxon, based on fossils only of teeth and these teeth have been found in the U.S. states of Arizona and New Mexico. The original report described it as a "probable ornithischian" and Heckert (2005) suggests that Krzyzanowskisaurus teeth have biostratigraphic utility as an index fossil of the St. Johnsian sub-LVF.
Desmatosuchus is an extinct genus of archosaur belonging to the Order Aetosauria. It lived during the Late Triassic.
Acaenasuchus is an extinct genus of pseudosuchian, endemic to what would be presently be known as Arizona during the Late Triassic, specifically during the Carnian and Norian stages of the Triassic. Acaenasuchus had a stratigraphic range of approximately 11.5 million years. Acaenasuchus is further categorized as one of the type fauna that belong to the Adamanian LVF, based on the fauna of the Blue Mesa Member of the Chinle Petrified Forest Formation of Arizona, where Acaenasuchus was initially discovered.
Smilosuchus is an extinct genus of leptosuchomorph parasuchid from the Late Triassic of North America.
The Cooper Canyon Formation is a geological formation of Norian age in Texas and New Mexico. It is one of several formations encompassed by the Dockum Group.
Sterling Nesbitt is an American paleontologist best known for his work on the origin and early evolutionary patterns of archosaurs. He is currently an associate professor at Virginia Tech in the Department of Geosciences.
Leptosuchus is an extinct genus of leptosuchomorph phytosaur with a complex taxonomical history. Fossils have been found from the Dockum Group and lower Chinle Formation outcropping in Texas, New Mexico, and Arizona, USA, and date back to the Carnian stage of the Late Triassic.
Machaeroprosopus is an extinct genus of mystriosuchin leptosuchomorph phytosaur from the Late Triassic of the southwestern United States. M. validus, once thought to be the type species of Machaeroprosopus, was named in 1916 on the basis of three complete skulls from Chinle Formation, Arizona. The skulls have been lost since the 1950s, and a line drawing in the original 1916 description is the only visual record of the specimen. Another species, M. andersoni, was named in 1922 from New Mexico, and the species M. adamanensis, M. gregorii, M. lithodendrorum, M. tenuis, and M. zunii were named in 1930. Most species have been reassigned to the genera Smilosuchus, Rutiodon, or Phytosaurus. Until recently, M. validus was considered to be the only species that has not been reassigned. Thus, Machaeroprosopus was considered to be a nomen dubium or "doubtful name" because of the lack of diagnostic specimens that can support its distinction from other phytosaur genera. However, a taxonomic revision of Machaeroprosopus, conducted by Parker et al. in 2013, revealed that UW 3807, the holotype of M. validus, is not the holotype of Machaeroprosopus, while the species Machaeroprosopus buceros, Machaeroprosopus being a replacement name, with a fixed type species, for Metarhinus, is the combinatio nova of the type species of the genu: Belodon buceros. Therefore, the name Pseudopalatus must be considered a junior synonym of Machaeroprosopus, and all species of the former must be reassigned to the latter. This revised taxonomy was already accepted in several studies, including Stocker and Butler (2013). Stocker and Butler (2013) also treated M. andersoni as a valid species, and not a junior synonym of Machaeroprosopus buceros as was previously suggested by Long and Murry (1995).
Parrishia is an extinct genus of sphenosuchian crocodylomorph known from the Late Triassic Chinle, Dockum, and Santa Rosa Formations in Arizona and New Mexico.
Aetosaurinae is one of the two main clades of aetosaurs, the other being Desmatosuchia. It is a stem-based taxon defined as all aetosaurs more closely related to Aetosaurus than Desmatosuchus. Aetosaurinae currently comprises Aetosaurus, similar forms such as Coahomasuchus and Stenomyti, and the widespread and successful aetosaur clade Typothoracinae.
The Colorado City Formation is a Late Triassic geologic formation in the Dockum Group of Texas, United States. It has previously been known as the Iatan Member, Colorado City Member or 'Pre-Tecovas Horizon'.
Scutarx is an extinct genus of Aetosauriformes, most commonly regarded by its species name Scutarx deltatylus. Scutarx lived around 230 million years ago during the Carnian and Norian stage of the Late Triassic. Scutarx are “medium sized” paramedian osteoderms belonging to the clade Aetosauria, a heavily armored and more herbivorous cousin of crocodiles.
Vivaron is a genus of rauisuchid known from the Late Triassic Chinle Formation in New Mexico. It is the second rauisuchid known from the southwestern United States, and it highlights the wide biogeographic range similar rauisuchid taxa occupied during the Late Triassic across Pangaea, despite the varied faunal assemblages at different latitudes.
Aetosauriformes is an extinct clade of early-diverging pseudosuchians. It includes the aetosaurs, a group of heavily armoured and at least partially herbivorous pseudosuchians, as well as the closely related genera Acaenasuchus, Euscolosuchus and Revueltosaurus.