Paleobiota of the Chinle Formation

The Chinle Formation is an extensive geological unit in the southwestern United States, preserving a very diverse fauna of Late Triassic (primarily Norian-age) animals and plants. This is a list of fossilized organisms recovered from the formation.

Amniotes

Archosauromorphs

Crurotarsans

Main article: Archosaurs of the Chinle Formation

Other Archosauromorphs

Non-crurotarsan Archosauromorphs of the Chinle Formation
Genus	Species	State	Member	Abundance	Notes	Images
Crosbysaurus [1]	C. harrisae [1]	Arizona Utah [1]	Blue Mesa [1]		An archosauriform represented only by teeth. [1] It is treated as indeterminate archosauriform remains by Irmis in 2005. [1] It was originally thought to be an ornithischian dinosaur.	Crosbysaurus Doswellia Trilophosaurus Vancleavea
Doswellia [2]	cf. D. kaltenbachi	Arizona	Blue Mesa [2]	Osteoderms, vertebrae, rib and hip fragments.	A doswelliid archosauriform.
Puercosuchus [3]	P. traverorum	Arizona	Blue Mesa	Two monodominant bonebeds.	A carnivorous azendohsaurid archosauromorph.
Rugarhynchos [4]	R. sixmilensis	New Mexico	Bluewater Creek	Skull and postcranial fragments.	A doswelliid archosauriform. Originally described as a species of Doswellia , [5] but subsequently transferred to a separate genus.
Syntomiprosopus [6]	S. sucherorum	Arizona	Blue Mesa or Sonsela [6]	Two to four individuals from a single quarry. [6]	A short-faced archosauriform, possibly an unusual early-diverging crocodylomorph. Appears convergent with some Late Cretaceous notosuchians. [6]
Tanystropheidae [7]	Indeterminate	New Mexico	Petrified Forest	Vertebrae	Moderately-sized tanystropheid vertebrae from the Hayden Quarry, likely representing a new taxon closely related to Langobardisaurus and Tanytrachelos . [7]
Tanytrachelos [8]	Indeterminate [8] / T. ahynis [7]	New Mexico Arizona [8]	Petrified Forest Mesa Redondo [8]	Cervical rib, [8] calcaneum, etc.	A small tanystropheid represented by several hundred fossil specimens.
Tecovasaurus [9]	T. murryi [9]	Arizona [9]	Mesa Redondo [9]		An unknown amniote represented by scattered teeth formerly believed to be from an ornithischian dinosaur. [9] Later discoveries of similar teeth in pseudosuchians meant that these could no longer be regarded as anything more specific than some kind of archosauriform. [9]
Trilophosaurus [10]	T. buettneri [11]	Arizona [11]	Blue Mesa [11]
	T. dornorum [12]	Arizona [12]	Sonsela [12]
	T. jacobsi [13]	Arizona [13]	Blue Mesa [13] Mesa Redondo [13]
	T. phasmalophos [14]	Arizona	Sonsela
Vancleavea [15]	V. campi [15]	Arizona [15] New Mexico	"Siltstone" [16] Petrified Forest [15] Sonsela [15] Blue Mesa [15]		A strange aquatic carnivorous archosauriform, represented by both articulated skeletons and scattered elements like osteoderms and vertebrae. [15]

Other amniotes

Miscellaneous amniotes of the Chinle Formation
Genus	Species	State	Member	Abundance	Notes	Images
Acallosuchus [17]	A. rectori [17]	Arizona [17]	Blue Mesa [17]	Known only from a partial skull. [17]	A strange neodiapsid whose bones were heavily ornamented "with subtriangular knobs... running the length of the bones." Even these ornamentations were ornamented "with additional grooves." It is too bizarre to be currently classified as anything more than a probable diapsid. [17]	Drepanosaurus Placerias
Ancistronychus [18]	A. paradoxus	Arizona	Sonsela	Manual unguals (hand claws).	A drepanosaur related to Drepanosaurus.
Avicranium [19]	A. renestoi	New Mexico	'Siltstone'	Skull and neck vertebrae.	A drepanosaur with a toothless skull and a flexible neck owing to the heterocoelous (saddle-shaped) articular surfaces of the vertebral centrae. The neural spines are anteroposteriorly short and strongly anterodorsally inclined. These features closely resemble those of Drepanosaurus .
Colognathus [20]	C. obscurus [20]	Arizona [20]	Blue Mesa [20]	Known only from a jaw fragment and some isolated teeth. [20]	Originally believed to be a fish, Colognathus was a strange amniote with distinctive fluted teeth. [20]
Dolabrosaurus	D. aquatilis	New Mexico	Petrified Forest	Articulated vertebral and limb material.	A drepanosaur.
Drepanosaurus [21]	D. unguicaudatus	New Mexico	Petrified Forest		A drepanosaur.
Kataigidodon [22]	K. venetus	Arizona	Blue Mesa	Partial dentaries.	A non-mammalian eucynodont.
Kraterokheirodon [23]	K. colberti [23]	Arizona [23]	Petrified Forest [23] Blue Mesa or Mesa Redondo [23]	Known only from two teeth. [23]	A possible cynodont. Although they share some similarities with cynodont teeth, the teeth of Kraterokheirodon are very distinctive and can't be confidently referred to a known amniote group. [23]
Palacrodon [24]	Indeterminate	Arizona	Blue Mesa	Jaw fragments.	A diapsid reptile of uncertain phylogenetic placement with unusual broadened teeth. More complete fossils of this genus from the Fremouw Formation of Antarctica suggest that it was an arboreal saurian. [25]
Placerias [26]	P. hesternus [26]	Arizona [26]	Blue Mesa [26] Mesa Redondo [26]	Known from several hundred remains, but very rare outside of the highly concentrated Placerias Quarry. [26]	A placeriine stahleckeriid dicynodont.
Skybalonyx [27]	S. skapter	Arizona	Blue Mesa	Manual unguals (hand claws).	A drepanosaur which likely had a burrowing lifestyle.
Uatchitodon [28]	U. schneideri [29]	Arizona [30]	Mesa Redondo [30]	Known in Chinle from only a single tooth. The presence of venom channels is consistent with other known Uatchitodon specimens, although the Chinle specimen's channels are unique in being "completely enclosed under the surface of the crown." [30]	A reptile of unknown affinities, probably a carnivorous archosauromorph with venomous capabilities.
Whitakersaurus [31]	W. bermani	New Mexico	"Siltstone"		A rhynchocephalian found at Ghost Ranch.
Indeterminate procolophonid [32]	Indeterminate	Utah

Amphibians

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon		Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Amphibians of the Chinle Formation
Genus	Species	State	Member	Abundance	Notes	Images
Apachesaurus [33]	A. gregorii [33]	Arizona [33]	Owl Rock [33] Petrified Forest [33] Blue Mesa [33]	Common in the Owl Rock and Petrified forest members. [33] Blue Mesa remains are fragmentary. [33]	A metoposaurid temnospondyl which is a junior synonym of Anaschisma	Anaschisma Funcusvermis
Anaschisma [34] [35]	A. browni [36] [35]	Arizona [36]	Owl Rock [36] Petrified Forest [36] Sonsela [36] Blue Mesa [36] Mesa Redondo [36]	A. browni is "possibly the most common tetrapod fossil in the lower Chinle", although its presence in the upper Chinle is "unclear". [36]	A metoposaurid temnospondyl. Koskinonodon was erected for the species "Buettneria" perfecta when it was discovered that the latter genus was preoccupied. Gee et al., 2017 then synonymised it with Anaschisma. [34] [35]
Chinlestegophis [37]	C. jenkinsi	Colorado			A temnospondyl in the group Stereospondyli, related to Rileymillerus from the Dockum Group of Texas. Possibly a stem-caecilian according to one analysis, [37] though others have disputed this finding
Funcusvermis [38]	F. gilmorei	Arizona	Blue Mesa	Numerous jaw fragments, as well as a referred maxillopalatine, vertebra, and femur	A stem-caecilian
Salientia indet. [39]	Indeterminate	Arizona	Sonsela Member Blue Mesa Member	Five bones: four ilia and a partial maxilla.	A stem-group frog. Might be more closely related to crown-group frogs (anurans) than to Early Triassic taxa Triadobatrachus and Czatkobatrachus .

Cartilaginous fish

Chondrichthyans of the Chinle Formation
Genus	Species	State	Stratigraphic position	Abundance	Notes	Images
Acrodus [40]	Indeterminate [40]	Arizona [40]	Petrified Forest [41] Sonsela [41] Mesa Redondo [40]	Only a single tooth is known. [40]	A hybodontiform shark.	Xenacanthus
Lonchidion [42]	L. humblei [42]	Arizona [42]	Blue Mesa [42] Mesa Redondo [42]		A hybodontiform shark. Lonchidion remains are common throughout the Carnian microvertebrate sites of the American southwest. [42]
Palaeoxyris	P. humblei [43]	Arizona	Blue Mesa	Several specimens	An egg capsule of a freshwater hybodont shark.
Phoebodus [44]	Indeterminate [44]	Arizona [44]	Mesa Redondo [44]	Only a single specimen has been recovered from the formation. [44]
Reticulodus [45]	R. synergus [45]	Arizona [45]	Owl Rock [45] Petrified Forest [45] Sonsela [45]		The crown of its tooth bears a "reticulating ornamentation on [its] occlusal surface[.]" [45] Reticulodus remains are common throughout the Norian microvertebrate sites of the American southwest. [45]
Xenacanthus [46]	X. moorei [46]	Arizona [46]	Blue Mesa [46] Mesa Redondo [46]	Common in the lower Chinle Formation's microvertebrate localities. [46]	A xenacanthiform shark.

Lobe-finned fish

Coelacanths

Actinistians (coelacanths) of the Chinle Formation
Genus	Species	State	Stratigraphic position	Abundance	Notes	Images
Chinlea [47]	Indeterminate [48]	Arizona [48]	Owl Rock [48] Blue Mesa [48]		An actinisitan (coelacanth).	Chinlea

Lungfish

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon		Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Dipnoans (lungfish) of the Chinle Formation
Genus	Species	State	Stratigraphic position	Abundance	Notes
Arganodus [49]	A. dorotheae [49]
	Indeterminate [49]	Arizona [49]	Petrified Forest [49] Sonsela [49] Blue Mesa [49] Mesa Redondo [49]	Arganodus toothplates are the most common fossil in the formation from a non-tetrapod. [49] They are evenly distributed across strata, although some individual localities have very high concentrations. [49]	Most Chinlean Arganodus fossils are isolated tooth plates. [49]
Ceratodus [49]	C. dorotheae [49]				Named by Case in 1921, in the 1980s it was referred to Arganodus. [49]

Ray-finned fish

Actinopterygians (ray-finned fish) of the Chinle Formation
Genus	Species	State	Stratigraphic position	Abundance	Notes	Images
Australosomus [50]	Indeterminate [51]	Arizona [51]	Mesa Redondo [51]	Known only from two vertebrae. [51]		Australosomus Saurichthys
Cionichthys [52]	C. dunklei	Colorado Utah			A member of Redfieldiiformes.
Hemicalypterus	H. weiri	Utah	Church Rock Member		A member of Dapediiformes. [53]
Lasalichthys [52] [54]	L. hillsi [52]	Utah [52]			A member of Redfieldiiformes.
	Indeterminate [55]	Arizona [55]	Petrified Forest [55] Blue Mesa [55] Mesa Redondo [55]		Represented by isolated scales. [55]
Lophionotus [56]	L. sanjuanensis [56]	Utah	Church Rock Member [56]		A member of Semionotiformes.
	L. chinleana [57]	Utah	Church Rock Member		A member of Semionotiformes.
Saurichthys [58]	Indeterminate	Arizona	Upper Blue Mesa Member		A member of Saurichthyiformes.
Synorichthys [52]	S. stewarti	Colorado Utah			A member of Redfieldiiformes.
Tanaocrossus [52]	T. kalliokoskii	Colorado			An enigmatic actinopterygian. [53]
Turseodus [52] [59]	T. dolorensis [52]	Colorado [52]			A member of Palaeonisciformes. [52]
	Indeterminate [60]	Arizona [60]	Owl Rock [60] Petrified Forest [60] Sonsela [60] Blue Mesa [60]	Common. [60]	Isolated scales from Chinle microvertebrate sites commonly have Turseodus-like ridges, however that feature is not unique to Turseodus and in 2005 Irmis advised researchers to regard them as indeterminate palaeoniscid remains. [60]

Plants

The Chinle Formation has a diverse flora of plant megafossils, though they are concentrated in only a few sites with suitable conditions. One of the most diverse floral communities is found near Fort Wingate, New Mexico. Paleobotanists have traditionally placed the Fort Wingate plant beds into the Monitor Butte Member, though more recently they are placed within the Bluewater Creek Formation, a subunit of the Chinle Formation first defined in 1989. ^[61] Some Fort Wingate plant fossils belong to the "Lake Ciniza beds", a localized patch of grey mudstone corresponding to an ancient lake. ^[62]

Another productive areas for plant fossils is Petrified Forest National Park in Arizona. Though petrified wood could be found through the entire stratigraphy of the park, most other plant fossils are exclusive to greenish mudstone layers adjacent to the Newspaper Rock sandstone bed in the Blue Mesa Member (formerly known as the "Lower Petrified Forest").

Conifers are the most common and diverse plants, including petrified wood and leafy branches from massive trees ( Araucarioxylon , Pagiophyllum ) as well as smaller shrubby forms ( Pelourdea ). Cycad and bennettitalean leaves and other remains make up a significant portion of the flora ( Zamites , Nilssoniopteris , Williamsonia , etc.). Ferns ( Cladophlebis , Phlebopteris , Clathropteris , Cynepteris, etc.) are abundant, with a range of growth habits including low shrubs, tree ferns, and palm-like fronds comparable to their modern relatives. Sphenophytes (horsetails: Neocalamites , Equistetites , Schizoneura , etc.) have low diversity but high abundance, and the largest Neocalamites fossils in the Chinle Formation could reach up to 6 meters (20 feet) in height. "Seed ferns" ( Chilbinia , Marcouia ?) ginkgophytes ( Baiera ), and small lycopods ( Chinlea , Selaginella ) were present but uncommon. The flora is rounded out by unusual low-growing gymnosperms such as Sanmiguelia (an angiosperm-like shrub), Dechellyia , and Dinophyton (possible relatives of Gnetales). ^{[63] [64]}

The floral composition of the Chinle Formation (and other parts of Late Triassic North America) seem to shift with changes in climate over time. The lowest parts of the Chinle, such as the Shinarump Conglomerate, are dominated by the bennettitalean Eoginkgoites alongside the first occurrence of other persistent plants such as Phlebopteris , Equisetites , and most common conifer species. Subsequent subunits (such as the Blue Mesa Member, Monitor Butte Member, and Bluewater Creek Formation) are much more diverse, with a wide array of humidity-adapted plants making up the typical Chinle flora. This second floral zone is characterized by Dinophyton , a common but enigmatic shrubby gymnosperm. Plant fossils are rare in the upper part of the Chinle Formation, which was presumably much drier than the lower part. In these later layers, by far the most common plant fossils belong to Sanmiguelia (an endemic of southwestern North America) alongside conifers and horsetails. ^{[65] [66]}

Gymnosperms

Gymnosperms of the Chinle Formation
Genus	Species	State	Stratigraphic position	Abundance	Notes	Images
Araucariorhiza [67]	A. joae	Arizona [67] [63]	"Petrified Forest" sensu lato [67] [63]	Uncommon	Roots of conifers similar to modern Araucariaceae. Possibly from the same plant as Araucarioxylon arizonicum .
Araucarioxylon [68]	A. arizonicum, A. sp. [69]	Arizona [63] Utah [69] New Mexico [70] [71]	"Siltstone" [63] [69] Petrified Forest [63] [65] Sonsela [65] Blue Mesa [65] Bluewater Creek (formerly Monitor Butte) [71] Agua Zarca Sandstone [70]	Abundant	Petrified trunks of large conifers possibly related to modern Araucariaceae. [72] State fossil of Arizona.	Araucarioxylon arizonicum
Araucarites	A. rudicula [73]	Arizona New Mexico	Blue Mesa Monitor Butte	Very rare	Large female conifer cones similar to those of araucariacean conifers.
Aricycas [74]	A. paulae	Arizona	Blue Mesa ( = "Lower Petrified Forest")	Moderately common	Pinnate cycad leaves
Baiera	B. arizonica	Arizona [63] New Mexico [75] [71]	Blue Mesa (= "Lower Petrified Forest") [63] [65] Bluewater Creek ( = "lower red member", formerly Monitor Butte) [75] [71]	Uncommon	Ginkgophyte leaves
Brachyphyllum	B. sp, B. hegewaldia [76]	Arizona [63] [76] New Mexico [70] [63] Utah [63]	Blue Mesa ( = "Lower Petrified Forest") [76] [63] Agua Zarca Sandstone [70] [63] Shinarump [63] Temple Mountain [63]	Uncommon	Conifer leaves and shoots
Carpolithus	C. chinleana	Arizona [63]	Blue Mesa (= "Lower Petrified Forest") [63] [65]	Rare	Seeds of uncertain affinities
Cephalotaxopsis	C. sp.	Arizona [63]			Conifer foliage
Charmorgia	C. dijolli	Arizona [63]	"Petrified Forest" sensu lato [63]	Rare	Short cycad stems
Chilbinia [77]	C. lichii	Arizona	Shinarump	Rare	An archaic seed fern with seeds and leaves more similar to Carboniferous-Permian seed ferns than to Triassic forms.
Creberanthus	C. bealeii [73]	Arizona	Blue Mesa [73] [65]	Very rare	Pollen-bearing cones similar to those of some extinct "pteridosperms" (seed ferns). Associated with Alisporites opii, a common palynomorph.
Dadoxylon	D. chaneyi	Arizona [63]	"Petrified Forest" sensu lato [63]	Very rare	Possible cordaitalean conifer wood.
Dechellyia [78]	D. gormanii	Arizona [78] [79] New Mexico [78] [71] [80]	Bluewater Creek (formerly Monitor Butte) [71] [80] Monitor Butte [78] [79] [80]	Uncommon	An enigmatic gymnosperm with narrow pinnate leaves, "clasping" leaf-like sporophylls, and winged seeds. Some leaves are affected by galls, similar to those formed by eriophyid mites in modern plants. [79] [80]
Dinophyton	D. spinosus [81]	Arizona [81] New Mexico [81] [62] [71]	Blue Mesa (= "Lower Petrified Forest") [81] [63] [65] Monitor Butte [81] Bluewater Creek ("Ciniza Lake beds", formerly Monitor Butte) [62] [71]	Common	An unusual gymnosperm combining needle-covered shoots with seed-bearing structures having the form of "pinwheel"-like clusters of tubular needles. Possibly related to Gnetales.
Elatocladus	E. puercoensis [80]	Arizona	Blue Mesa	Uncommon	Leaf-bearing conifer shoots
Eoginkgoites	E. sp., E. davidsonii [82]	Arizona [82] Utah [82]	Shinarump [82]	Common within its small stratigraphic range	A bennettitalean with palm-shaped fronds similar to some ginkgo species.
Ginkgoites	G. sp.	Arizona [65]	Blue Mesa [65]	Rare	A ginkgophyte
Lindleycladus	L. arizonicus [80]	Arizona	Blue Mesa	Uncommon	Conifer shoots bearing Podozamites leaves. Formerly known as Podozamites arizonicus.
Lyssoxylon	L. grigsbyi	Arizona [63] New Mexico [71]	"Petrified Forest" sensu lato [63] Bluewater Creek (formerly Monitor Butte) [71]	Rare	Cycad trunks
Marcouia	M. neuropteroides	Arizona [63] New Mexico [71] [79]	Blue Mesa (= "Lower Petrified Forest") [63] [65] Bluewater Creek (formerly Monitor Butte) [71] Monitor Butte [79]	Uncommon	Fern-like gymnosperm leaves of uncertain affinities. Some leaves are affected by feeding traces including crescent-shaped marginal incisions and oval-shaped holes. These were probably produced by beetles or orthopterans (grasshoppers and kin). [79]
Masculostrobus	M. clathratus [78]	Arizona	Monitor Butte	Locally common	A pollen-bearing cone associated with lattice-shaped Equisetosporites chinleana pollen.
Nilssoniopteris	N. ciniza [62] [71]	Arizona [63] New Mexico [75] [62] [71] Utah [63]	"Petrified Forest" sensu lato [63] Bluewater Creek ("Ciniza Lake beds", formerly Monitor Butte) [75] [62] [71] Shinarump [63]	Uncommon	Large smooth-edged bennettitalean leaves, some of which were originally identified as Macrotaeniopteris magnifola.
Otozamites	O. macombii	New Mexico [70]	Agua Zarca Sandstone [70]	Uncommon	Bennettitalean leaves
Pagiophyllum	P. sp., P. duttonia, [62] P. navajoensis, [62] P. readiana, [62] P. zuniana, [62] P. simpsonii [83]	Arizona [83] Colorado [69] New Mexico [70] [62] [71]	"Middle Dolores" [69] "Petrified Forest" sensu lato [63] Sonsela [83] [65] Bluewater Creek ("Ciniza Lake beds", formerly Monitor Butte) [62] [71] Agua Zarca Sandstone [70]	Locally very common	Conifer leaves
Palissya	P. sp., P. diffusa, P. sphenolepis	New Mexico [63] Utah [63]	Monitor Butte [63] Agua Zarca Sandstone [63] Shinarump [63]	Uncommon	Conifer leaves and shoots
Pelourdea	P. poleoensis	Utah [69] Colorado [69] New Mexico [62] [71]	"Siltstone" [69] "Middle Dolores" [69] Poleo [63] Bluewater Creek ("Ciniza Lake beds", formerly Monitor Butte) [62] [71] Shinarump [63]	Common	A shrubby conifer with strap-like leaves arranged around a single narrow stem.
Podozamites	P. sp., P. emmonsi, P. lanceolatus	Arizona [63] Utah [63]	Blue Mesa (= "Lower Petrified Forest") [63] [65] Shinarump [63] Temple Mountain [63]	Uncommon	Conifer leafs and shoots
Pramelreuthia	P. dubielli, [84] P. yazzi [84]	Arizona New Mexico Utah	Blue Mesa [65] Monitor Butte	Rare	Gymnosperm microsporophylls (pollen-bearing organs)
Pterophyllum	P. sp., P. braunianum	Arizona [63] Utah [63]	Shinarump [63]	Uncommon	Bennettitalean leaves
Samaropsis	S. sp., S. puerca	Arizona [63]	Blue Mesa (= "Lower Petrified Forest") [63] [65]	Uncommon	Seeds of uncertain affinities
Sanmiguelia	S. lewisii	Arizona [69] [85] Utah [86] [69] [85] Colorado [69]	Rock Point [69] Owl Rock [86] [69] "Middle Dolores" [69] Petrified Forest [85] Sonsela [85] Blue Mesa [85]	Common	An unusual palm-like shrub, controversially suggested to be a relative or predecessor of angiosperms (flowering plants).
Schilderia	S. adamanica	Arizona [63]	Petrified Forest [63] [65]	Rare	Petrified stems of a conifer or gnetalean [72]
Williamsonia	W. nizhonii [87]	New Mexico [75] [87] [71]	Bluewater Creek ( = "lower red member", formerly Monitor Butte) [75] [87] [71]	Rare	A bennettitalean "flower" (female cone)
Woodworthia	W. arizonica	Arizona [63]	Petrified Forest [63] [65]	Uncommon	Stems and petrified wood of a conifer related to A. arizonicum. [72] [88]
Zamites	Z. sp., Z. occidentalis, Z. powellii	Arizona [89] [79] Utah [90] [79] New Mexico [75] [70] [62] [71]	Church Rock? [90] Owl Rock? [90] Blue Mesa (= "Lower Petrified Forest") [63] [65] Sonsela [89] [65] Bluewater Creek ("Ciniza Lake beds", formerly Monitor Butte) [75] [62] [71] Monitor Butte [79] Agua Zarca [70] Shinarump [79]	Very common	Bennettitalean leaves. Some leaves exhibit feeding traces, including sharp diagonal marginal traces and slot-like non-marginal traces. These traces were probably left by beetles. [79] Zamites powellii was formerly known as Otozamites powellii.

Ferns

Ferns of the Chinle Formation
Genus	Species	State	Stratigraphic position	Abundance	Notes	Images
Apachea	A. arizonica	Arizona [63]	"Petrified Forest" [63]	Rare	Dipteridacean fern leaves, possibly damaged specimens of Clathropteris walkeri. [91]
Cladophlebis	C. daughertyi, [91] C. subfalcata, C. yazzia, [76] C. sp.	Arizona [91] [76] [63] New Mexico [75] [91] [71] [63] Utah [63]	Blue Mesa (= "Lower Petrified Forest") [91] [76] [65] Bluewater Creek (= "lower red member", formerly Monitor Butte) [75] [91] [71] Monitor Butte [63] Shinarump [63]	Very common	Large sterile fern leaves of uncertain affinities
Clathropteris	C. walkeri	Arizona [91] [63] New Mexico [75] [91] [71] [63]	Blue Mesa (= "Lower Petrified Forest") [91] [65] Bluewater Creek (= "lower red member", formerly Monitor Butte) [75] [91] [71] [63]	Common	Dipteridacean fern leaves
Cynepteris [91]	C. sp., C. lasiophora [91]	Utah [90] Arizona [91] [79] New Mexico [91] [71] [79]	Church Rock? [90] Owl Rock? [90] Blue Mesa (= "Lower Petrified Forest") [91] [79] Bluewater Creek (= "lower red member", formerly Monitor Butte) [91] [71] Monitor Butte [79]	Common	Bipinnate fern leaves previously identified as Lonchopteris . Some leaves are affected by crescent-shaped marginal feeding traces, probably from beetles or orthopterans (grasshoppers and kin). [79]
Itopsidema [92]	I. vancleavi	Arizona	Sonsela	Uncommon	Stems of large osmundacean tree ferns. Some stems show evidence of being bored by oribatid mites. [93]
Phlebopteris	P. smithii, P. utensis	Arizona [91] [94] New Mexico [75] [91] [62] [71] Utah [94]	Blue Mesa (= "Lower Petrified Forest") [94] [65] Bluewater Creek ( = "lower red member" / "Ciniza Lake beds", formerly Monitor Butte) [75] [91] [62] [71] Shinarump [94]	Common	Matoniacean fern leaves with a palmate structure
Sphenopteris	S. arizonica	Arizona [63] [95]	Blue Mesa (= "Lower Petrified Forest") [63] [65]	Uncommon	Sterile fern leaves of uncertain affinities. Some show evidence for marginal damage by insect feeding traces. [95]
Todites	T. fragilis	Arizona [91] New Mexico [75] [91] [71]	Blue Mesa (= "Lower Petrified Forest") [91] Bluewater Creek (="lower red member", formerly Monitor Butte) [75] [91] [71]	Uncommon	Osmundacean fern leaves
Wingatea [91]	W. plumosa	Arizona [91] New Mexico [91] [71]	Blue Mesa (= "Lower Petrified Forest") [91] [65] Bluewater Creek (="lower red member", formerly Monitor Butte) [91] [71]	Rare	Gleicheniacean fern leaves previously named as a species of Coniopteris .

Other plants

Other plants of the Chinle Formation
Genus	Species	State	Stratigraphic position	Abundance	Notes	Images
Chinlea	C. sp., C. campii	Arizona [96] [63]	"Petrified Forest" sensu lato [63]	Uncommon	Small lycopod stems similar to shrubby forms such as Pleuromeia . [96]
Equisetites	E. sp., E. bradyi	Arizona [63] Utah [63] New Mexico [62] [71]	Blue Mesa (= "Lower Petrified Forest") [63] [65] Bluewater Creek ("Ciniza Lake beds", formerly Monitor Butte) [62] [71] Shinarump [63]	Common	A horsetail indistinguishable from modern Equisetum .
Equisetocalis [97]	E. muirii	Arizona	Blue Mesa	Concentrated in a narrow stratigraphic range (the Newspaper Rock sandstone bed)	Small three-dimensional horsetail stems preserved in pyrite nodules.
Isoetites	I. circularis	Arizona [63]	"Petrified Forest" sensu lato [63]	Rare	Quillwort shoots
Lycostrobus	L. chinleana	Arizona [63]	Blue Mesa (= "Lower Petrified Forest") [63] [65]	Rare	Horsetail cones sometimes placed in the genus Equicalostrobus
Neocalamites	N. sp., N. virginiensis	Arizona [63] Utah [90] [69] Colorado [98] [69] New Mexico [75] [62]	"Siltstone" [69] Church Rock? [90] Owl Rock? [90] "Petrified Forest" sensu lato [63] Bluewater Creek ( = "lower red member" / "Ciniza Lake beds", formerly Monitor Butte) [75] [62]	Common	Stems and leaves of a giant sphenophyte (horsetail)
Schizoneura	S. harrisii	New Mexico [71]	Bluewater Creek (formerly Monitor Butte) [71]	Uncommon	Stems and leaves of a large sphenophyte (horsetail) more typical of Southern Hemisphere plant assemblages.
Selaginella	S. anasazia [78]	Arizona	Monitor Butte	Uncommon	A low-growing lycopsid similar to modern broad-leaved species of Selaginella, such as S. kraussiana

Arthropods

Arthropods of the Chinle Formation
Genus	Species	State	Stratigraphic position	Abundance	Notes	Images
Enoplocytia	E. porteri [99]	Arizona	Sonsela?	Single well-preserved fossil	An erymid crayfish
Paleoscolytus [100]	P. divergus	Arizona	Sonsela		A species of bark beetle, evidenced by trace fossils of tracks made on A. arizonicum specimens.

References

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