Metapelma archetypon Temporal range: | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hymenoptera |
Family: | Metapelmatidae |
Genus: | Metapelma |
Species: | †M. archetypon |
Binomial name | |
†Metapelma archetypon Gibson, 2009 | |
Metapelma archetypon is an extinct species of parasitic wasp in the Metapelmatidae genus Metapelma . [1] The species is solely known from the Early Eocene [2] Baltic amber deposits in the Baltic Sea region of Europe. [1] Of the thirty seven described species in the genus Metapelma, M. archetypon is the only species known from the fossil record. [1]
Metapelma archetypon is known only from one fossil, the holotype, number "AMNH BaJWJ-407", which is a single female specimen preserved, along with a spider, in a nearly rectangular amber block 13 by 11 millimetres (0.51 by 0.43 in) in size. The block is currently residing in the American Museum of Natural History paleoentomology collections in New York City, USA. [1] M. archetypon was first studied by Gary A. P. Gibson with his 2009 type description being published in the journal ZooKeys . [1] The specific epithet "archetypon" was designated by Gary Gibson from the Greek ἀρχέτυπον (archetypon) which translates as "original" or "model". This is in reference to the similarity of the head and hind leg structure similarity between the extinct and extant species. [1]
The holotype of Metapelma archetypon is 7.7 millimetres (0.30 in) in length when the ovipositor is excluded. Several areas of the female are obscured or missing, with the dorsal mesosomal and part of the gastral structures covered by a white substance. The terminal sections of both antenna and three of the legs are missing and both hind tarsi have the apical three segments detached but still present. Also missing is the terminal section of the ovipositor sheaths. [1] The forewings are hyaline in coloration. While M. archetypon has a deeply divided upper and lower mesepimeron separating the acropleuron and metapleuron, a feature found only in Metapelma, the shapes of the hind legs and head are closer to the general morphology of the subfamily Neanastatinae. The basal Neanastatinae groundplan is thought to have been modified in the modern genera in the subfamily. Although not confirmed shape of the ovipositor and relationship in the genus Metapelma suggests M. archetypon was parasitic on wood-boring beetles. [1]
The Mymarommatidae, sometimes referred to as false fairy wasps, are a very small family of microscopic parasitic wasps. Only about half of the known species are living taxa, but they are found worldwide.
The Stephanidae, sometimes called crown wasps, are a family of parasitoid wasps. They are the only living members of the superfamily Stephanoidea. Stephanidae has at least 345 living species in 11 genera. The family is considered cosmopolitan in distribution, with the highest species concentrations in subtropical and moderate climate zones. Stephanidae also contain four extinct genera described from both compression fossils and inclusions in amber.
Baltic amber or succinite is amber from the Baltic region, home of its largest known deposits. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that this forested region provided the resin for more than 100,000 tons of amber. Today, more than 90% of the world's amber comes from Kaliningrad Oblast of Russia. It is a major source of income for the region; the local Kaliningrad Amber Combine extracted 250 tonnes of it in 2014 and 400 tonnes in 2015.
Prionomyrmex is an extinct genus of bulldog ants in the subfamily Myrmeciinae of the family Formicidae. It was first described by Gustav Mayr in 1868, after he collected a holotype worker of P. longiceps in Baltic amber. Three species are currently described, characterised by their long mandibles, slender bodies and large size. These ants are known from the Eocene and Late Oligocene, with fossil specimens only found around Europe. It is suggested that these ants preferred to live in jungles, with one species assumed to be an arboreal nesting species. These ants had a powerful stinger that was used to subdue prey. In 2000, it was suggested by Cesare Baroni Urbani that the living species Nothomyrmecia macrops and a species he described both belonged to Prionomyrmex, but this proposal has not been widely accepted by the entomological community. Instead, scientists still classify the two genera distinctive from each other, making Nothomyrmecia a valid genus.
Electrinocellia is an extinct monotypic genus of snakefly in the family Inocelliidae containing the single species Electrinocellia peculiaris and known from Eocene aged Baltic amber.
Styporaphidia is a genus of snakefly, belonging to the extinct family Mesoraphidiidae, containing up to three species, the type species Styporaphidia magia, Styporaphidia willmanni and tentatively Styporaphidia? hispanica. The genus was named from the Greek stypos meaning "stem" or "stump" and Raphidia, the type genus for, and most often used as, a stem for generic names in the order Raphidioptera. The species name of S. magia is from the Greek word mageia meaning "magic" while the species name for S.? hispanica is from the Latin Hispania meaning "Spain" in reference to the type locality of the species.
Araneagryllus is an extinct monotypic genus of cricket in the subfamily Phalangopsinae with the sole species Araneagryllus dylani. The fossil was recovered in the Dominican Republic from early Miocene Burdigalian stage Dominican amber deposits on the island of Hispaniola. Araneagryllus is the first Phalangopsinae cricket to be described from the fossil record.
Aspidopleura is an extinct monotypic genus of parasitic wasp in the Eupelmidae subfamily Neanastatinae and at present, it contains the single species Aspidopleura baltica. The genus is solely known from the Early Eocene Baltic amber deposits in the Baltic Sea region of Europe.
Brevivulva is an extinct genus of parasitic wasp in the Eupelmidae subfamily Neanastatinae and contains the single species Brevivulva electroma. The genus is solely known from the Early Eocene Baltic amber deposits in the Baltic Sea region of Europe.
Neanaperiallus is an extinct monotypic genus of parasitic wasp in the Eupelmidae subfamily Neanastatinae containing only a single species, Neanaperiallus masneri. The genus is solely known from the Early Eocene Baltic amber deposits in the Baltic Sea region of Europe.
Glisachaemus is an extinct monotypic genus of planthopper in the Cixiidae subfamily Cixiinae and at present, it contains the single species Glisachaemus jonasdamzeni. The genus is solely known from the Early Eocene Baltic amber deposits in the Baltic Sea region of Europe.
Avitomyrmex is an extinct genus of bulldog ants in the subfamily Myrmeciinae which contains three described species. The genus was described in 2006 from Ypresian stage deposits of British Columbia, Canada. Almost all the specimens collected are queens, with an exception of a single fossilised worker. These ants are large, and the eyes are also large and well-developed; a sting is present in one species. The behaviour of these ants may have been similar to extant Myrmeciinae ants, such as foraging solitarily for arthropod prey and never leaving pheromone trails to food sources. Avitomyrmex has not been assigned to any tribe, instead generally being regarded as incertae sedis within Myrmeciinae. However, its identity as an ant has been challenged, although it is undoubtedly a hymenopteran insect.
Macabeemyrma is an extinct genus of bulldog ants in the subfamily Myrmeciinae containing the single species Macabeemyrma ovata, described in 2006 from Ypresian stage deposits of British Columbia, Canada. Only a single specimen is known; a holotype queen found preserved as a compression fossil. The specimen had no wings and small portions of its legs and eyes were faintly preserved. It was a large ant, reaching 25 millimetres (0.98 in) in length. This ants' behaviour would have been similar to that of extant Myrmeciinae ants, such as foraging singly in search for arthropod prey and nesting in soil or in trees. Macabeemyrma shows similarities to extinct ants in the genus Ypresiomyrma, and to the living Nothomyrmecia macrops, but has not been conclusively assigned to any tribe, instead generally regarded as incertae sedis within Myrmeciinae. However, the sole specimen lacks definitive traits, and its classification in Myrmeciinae, and even its identity as an ant, has been challenged.
Alavaraphidia is an extinct genus of snakefly in the family Mesoraphidiidae. The genus is solely known from an Early Cretaceous, Albian age, fossil amber found in Spain. Currently, the genus comprises a single species, Alavaraphidia imperterrita.
Arostropsis is an extinct genus of broad-nosed weevil in the beetle family Curculionidae known from an Upper Eocene fossil found in Europe. The genus contains a single described species, Arostropsis groehni.
Protostephanus is an extinct genus of crown wasp in the Hymenoptera family Stephanidae known from an Eocene fossil found in the United States of America. The genus contains a single described species, Protostephanus ashmeadi placed in the stephanid subfamily Stephaninae.
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