Molaria Temporal range: | |
---|---|
M. spinifera lectotype | |
Life restoration of M. spinifera | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
(unranked): | † Artiopoda |
Genus: | † Molaria Walcott, 1912 [1] |
Type species | |
Molaria spinifera Walcott, 1912 | |
Other species | |
Molaria steiniPeel, 2017 |
Molaria is a genus of Cambrian arthropod, the type species M. spinifera is known from the Middle Cambrian Burgess Shale. 144 specimens of Molaria are known from the Greater Phyllopod bed, where they comprise 0.27% of the community. [2] A second species M. steini was described from the Sirius Passet in Greenland in 2017. [3]
The genus name derives from “Molar”, the name of a mountain peak east of the Valley of the Ten Peaks in Alberta, Canada, [1] which in turn was so named by James Hector in 1859 because the shape of the mountain resembles a molar tooth. The specific name of the type species, spinifera, comes from Latin spinifer “spine-bearing.” [4]
The body of Molaria consisted of a head shield (cephalon), a trunk consisting of eight sections (tergites), and a telson, which included a short ventral spine and a long posterior spine. Three pairs of legs were beneath the cephalon and another eight pairs were attached to the trunk. Eyes were lacking, but a pair of short antennae was present on the cephalon. Specimens of Molaria ranged from 8 to 26 mm in length from cephalon to telson, with the posterior spine slightly longer than the body length. [5]
Molaria was superficially similar to Habelia , another Burgess Shale arthropod with a long tail spine, but which possessed 12 trunk tergites. [5] It is currently considered a member of Artiopoda. [3]
Opabinia regalis is an extinct, stem group arthropod found in the Middle Cambrian Burgess Shale Lagerstätte of British Columbia. Opabinia was a soft-bodied animal, measuring up to 7 cm in body length, and its segmented trunk had flaps along the sides and a fan-shaped tail. The head shows unusual features: five eyes, a mouth under the head and facing backwards, and a clawed proboscis that probably passed food to the mouth. Opabinia probably lived on the seafloor, using the proboscis to seek out small, soft food. Fewer than twenty good specimens have been described; 3 specimens of Opabinia are known from the Greater Phyllopod bed, where they constitute less than 0.1% of the community.
Marrella is an extinct genus of marrellomorph arthropod known from the Middle Cambrian of North America and Asia. It is the most common animal represented in the Burgess Shale of British Columbia, Canada, with tens of thousands of specimens collected. Much rarer remains are also known from deposits in China.
Sidneyia is an extinct marine arthropod known from fossils found from the Early to the Mid Cambrian of China and the Mid Cambrian Burgess Shale of British Columbia, Canada.
Yohoia is an extinct genus of megacheiran arthropod from the Cambrian period that has been found as fossils in the Burgess Shale formation of British Columbia, Canada. The type species, Yohoia tenuis, was described in 1912 by Walcott, who considered it an anostracan crustacean. 711 specimens of Yohoia are known from the Greater Phyllopod bed, where they comprise 1.35% of the community. In 2015, Conway Morris et al. reported another species, Y. utahana, from the Marjum Formation, Utah.
Canadaspis is an extinct genus of bivalved Cambrian arthropod, known from North America and China. They are thought to have been benthic feeders that moved mainly by walking and possibly used its biramous appendages to stir mud in search of food. They have been placed within the Hymenocarina, which includes other bivalved Cambrian arthropods.
Cheloniellida is a taxon of extinct Paleozoic arthropods. As of 2018, 7 monotypic genera of cheloniellids had been formally described, whose fossils are found in marine strata ranging from Ordovician to Devonian in age. Cheloniellida has a controversial phylogenetic position, with previous studies associated it as either a member or relative of various fossil and extant arthropod taxa. It was later accepted as a member of Vicissicaudata within Artiopoda.
Waptia is an extinct genus of arthropod from the Middle Cambrian of North America. It grew to a length of 6.65 cm (3 in), and had a large bivalved carapace and a segmented body terminating into a pair of tail flaps. It was an active swimmer and likely a predator of soft-bodied prey. It is also one of the oldest animals with direct evidence of brood care. Waptia fieldensis is the only species classified under the genus Waptia, and is known from the Burgess Shale Lagerstätte of British Columbia, Canada. Specimens of Waptia are also known from the Spence Shale of Utah, United States.
Olenoides was a trilobite from the Cambrian period. Its fossils are found well-preserved in the Burgess Shale in Canada. It grew up to 10 cm long.
Buenellus higginsi is an average size trilobite, which lived during the Lower Cambrian period, in what is now North-West Greenland. It is a prominent member of the Sirius Passet fauna. Buenellus higginsi is the only known species in the genus Buenellus.
Habelia is a genus of extinct arthropod from the Middle Cambrian, thought to be one of the earliest known relatives of chelicerates. Its fossils have been found in the Burgess Shale in British Columbia, Canada. Fifty-four specimens of Habelia are known from the Greater Phyllopod bed, where they comprise 0.1% of the community.
A number of assemblages bear fossil assemblages similar in character to that of the Burgess Shale. While many are also preserved in a similar fashion to the Burgess Shale, the term "Burgess Shale-type fauna" covers assemblages based on taxonomic criteria only.
Sarotrocercus is a small Cambrian arthropod known from Burgess shale, reaching a centimetre or two in length (0.39–0.79 in). Sarotrocercus is only known from 7 specimens. It may lie in the arthropod crown group, and a recent study has revised some points of its original description.
Plenocaris plena is a genus of extinct bivalved hymenocarine arthropod that lived in the Cambrian aged Burgess Shale and Chengjiang. Originally described as a species of Yohoia by Walcott in 1912, it was placed into its own genus in 1974.
Isoxys is a genus of extinct bivalved Cambrian arthropod; the various species of which are thought to have been freely swimming predators. It had a pair of large spherical eyes, and two large frontal appendages used to grasp prey.
Emeraldella is a genus of arthropod known from the Middle Cambrian of North America. The type species E. brocki was described in 1912 from the Burgess Shale. 21 specimens of Emeraldella are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community. A re-study on the species was done in 2012. A second species E. brutoni is known from the Wheeler Shale, which was described in 2011. An additional specimen of E. brutoni was described in 2019, which revealed more of the anatomy. It has been placed as a basal member of the clade Vicissicaudata within Artiopoda, a group of arthropods containing trilobites and their relatives.
Mollisonia is an extinct genus of Cambrian arthropod. 4 species had been described from North America and China. Studies suggest it is a basal member of Chelicerata, a group which includes horseshoe crabs and arachnids.
Kiisortoqia soperi is an extinct species of arthropod from the Early Cambrian Sirius Passet Lagerstätte in Greenland. While it had a superficially trilobite-like bodyform, it also possessed large frontal appendages similar to those of radiodonts.
Buenaspis is a genus of small nektaspid arthropod, that lived during the early Cambrian period. Fossil remains of Buenaspis were collected from the Lower Cambrian Sirius Passet Lagerstätte of North Greenland. Buenaspis looks like a soft eyeless trilobite. It has a headshield slightly larger than the tailshield (pygidium), and in between them six thoracic body segments (somites). The genus is monotypic, its sole species being Buenaspis forteyi.
Squamacula is an extinct artiopodan arthropod from the Cambrian Series 2. The type species S. clypeata was described in 1997 from the Chengjiang biota of Yunnan, China. At the time of description there were only two known specimens of S. clypeata, but now there are at least six known specimens. In 2012 a second species S. buckorum was described from the Emu Bay Shale of Australia.
Bushizheia is an extinct genus of arthropods known from a single specimen found in the Early Cambrian Maotianshan Shales Lagerstätte in China. The genus contains a single species, Bushizheia yangi. While it had a superficially trilobite-like bodyform, it also possessed large frontal appendages similar to those of radiodonts. It is morphologically similar to Kiisortoqia to which it is probably closely related.