Kangacaris Temporal range: | |
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Kangacaris zhangi | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Order: | † Nektaspida |
Family: | † Emucarididae |
Genus: | † Kangacaris Paterson et al., 2010 |
Species | |
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Kangacaris is an extinct genus of soft-shelled trilobite-like arthropod of the nektaspid order from the Lower Cambrian (Botomian). K. zhangi is known from South Australia, [1] and K. shui from South-West China. [2]
The generic name is derived from "Kanga", a contraction of Kangaroo Island, and Latin caris ("shrimp"). The specific name of the type species honors the Chinese palaeontologist Xingliang Zhang. [1]
The dorsal exoskeleton of Kangacaris is inverted egg-shaped, about 3 centimetres (1.2 in) long and 2 cm (0.79 in) wide. The axis is ⅓× as wide as the body and only slightly raised. The semi-circular headshield (or cephalon) is about ⅔× as long as the tailshield (pygidium), and in between them three short thoracic body segments (somites). The lateral margin of the pygidium is progressively angling towards the axis, ending in the slightest hint of a point. Twelve to thirteen furrows are most distinct on the axis, and becoming indiscernible near the border. The border of the cephalon is about ¼× as long (axially) as the thoracal somites, and the border of the pygidium is about ½× as long as the somites. This border has about 17 regularly spaced ridges perpendicular to the rim. [1] K. shui has a better defined and narrower axis, more arched posterior margin of the cephalic shield, more posterolaterally deflected pleural areas of the thoracic segments, and smaller pygidium than K. zhangi. [2]
Kangacaris differs from Emucaris fava in having three thorax segments, while Emucaris has four. When disregarding the border, Kangacaris has an inverted egg-shape, while the pygidium of Emucaris is a triangle with a rounded termination. The axis of the pygidium of Kangacaris is ⅓ of the width of the body and clearly segmented, while in Emucaris it is one fifth and has a pattern of polygons of approximately equal area. The axis of the pygidium of Emucaris terminates in a spine that ends at the outer rim of the border, while Kangacaris lacks a terminal spine.
Fossils of K. zhangi were collected from the Emu Bay Shale of Kangaroo Island, South Australia. Remains of K. shui have been found at the Maotianshan Shales, Yunnan, South-West China.
Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are Fallotaspis, and Lemdadella, both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian.
Emuellidae are a small family of trilobites, a group of extinct marine arthropods, that lived during the late Lower Cambrian of the East Gondwana supercontinent, in what are today South-Australia and Antarctica. Emuellidae can be recognized among trilobites in having a set of unique features. The headshield or cephalon has large genal spines reaching back as far as the 3rd to 6th segment of the thorax. The eye-ridges contact the back of the frontal lobe of the glabella and extend laterally and backwards, roughly parallel to the frontal and lateral rim of the cephalon. There are small, clearly incised pits at the junction between the eye-ridge and the frontal lobe of the cephalic axis. The thorax reaches its greatest width at the 6th segment. The frontal part or prothorax consists of 6 segments, with number 5 and 6 fused, and the 6th carrying very large trailing spines. The rear part or opistothorax consists of a variable but extremely large number of segments.
Redlichiina is a suborder of the order Redlichiida of Trilobites. The suborder contains three superfamilies: Emuelloidea, Redlichioidea and Paradoxidoidea. These trilobites are some of the oldest trilobites known. They originated at the beginning of the Cambrian Period and disappeared at the end of the middle Cambrian.
Redlichia is a genus of redlichiid trilobite in the family Redlichiidae, with large to very large species. Fossils of various species are found in Lower Cambrian (Toyonian)-aged marine strata from China, Korea, Pakistan, the Himalayas, Iran, Spain, southern Siberia, and Antarctica, and from Middle Cambrian (Ordian)-aged marine strata of Australia.
The Emu Bay Shale is a geological formation in Emu Bay, South Australia, containing a major Konservat-Lagerstätte. It is one of two in the world containing Redlichiidan trilobites. The Emu Bay Shale is dated as Cambrian Series 2, Stage 4, correlated with the upper Botomian Stage of the Lower Cambrian.
Waptia fieldensis is an extinct species of arthropod from the Middle Cambrian Burgess Shale Lagerstätte of Canada. It grew to a length of about 8 cm (3 in) and resembled modern shrimp in both morphology and habit. It had a large bivalved carapace and a segmented body terminating into a pair of tail flaps. It was an active swimmer, feeding on organic particles it gathered from the seafloor substrate. It is also one of the oldest animals with direct evidence of brood care.
Resserops is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the middle of the Atdabanian or the early part of the Botomian stage, which lasted from approximately 524 to 518.5 million years ago. This faunal stage was part of the Cambrian Period. It has been found in Spain and southern Morocco. It can be recognised by the sabre-like spines of the headshield that are a smooth continuation of the frontal edge, and the enlarged spines on the 9th segment of the thorax.
Holyoakia is a genus of very small trilobites of the family Dorypygidae, from the late Lower Cambrian of South Australia and Antarctica.
Yunnanocephalus is a genus of ptychopariid trilobite. It lived during the late Atdabanian and Botomian stages, in what are currently Antarctica, Australia and China. It was a "moderately common" member of the Chengjiang Fauna. Yunnanocephalus is the only genus currently assigned to the Yunnanocephalidae family.
Tsunyidiscus is a trilobite belonging to the Suborder Eodiscina. Tsunyidiscus appeared near the end of the Lower Cambrian, during the late Atdabanian stage of geologic time and some collections suggest it may have survived into the Botomian. The genus is very small, oculate and isopypous with a narrow dome-shaped glabella and a narrow bullet-shaped pygidial axis. Thorax consists of three segments. Tsunyidiscus is the only genus currently attributed to the family Tsunyidiscidae.
Panlongia was a small-sized marine arthropod, with an oval-shaped non-calcified exoskeleton. Both the head shield and the tail shield are semi-circular. In between the cephalon and pygidium are four thoracic body segments (somites). The cephalon occupies approximately ⅓ of the body length, the thorax ¼ and pygidium about 45%. Panlongia lived during the late Lower Cambrian (Botomian) in what is today South China. In Panlongia spinosa the edge of the exoskeleton carries several small sawtooth-like spines, that are absent in P. tetranodusa.
Aaveqaspis is a genus of small marine arthropods of unclear affiliation, that lived during the early Cambrian period. Fossil remains of Aaveqaspis were collected from the Lower Cambrian Sirius Passet fossil-Lagerstätte of North Greenland. Aaveqaspis looks like a soft eyeless trilobite with a weakly defined axis, a headshield with stubby genal spines, 5 thorax segments also ending in stubby genal spines, and a tailshield (pygidium) with a pair of massive tusk-like spines, and two smaller spines near the end of the axis. The only species presently known is A. inesoni.
Buenaspis is a genus of small marine arthropods in the family Liwiidae, that lived during the early Cambrian period. Fossil remains of Buenaspis were collected from the Lower Cambrian Sirius Passet Lagerstätte of North Greenland. Buenaspis looks like a soft eyeless trilobite. It has a headshield slightly larger than the tailshield (pygidium), and in between them six thoracic body segments (somites). The genus is monotypic, its sole species being Buenaspis forteyi.
Emucarididae is an extinct family of soft-shelled trilobite-like arthropods (nektaspids) from the Lower Cambrian of South Australia and South China. It contains only two genera – Emucaris and Kangacaris. Two species were described in 2010 from specimens recovered from Emu Bay Shale Lagerstätte, one species in 2012 from the Maotianshan Shales. It is classified under the order Nektaspida, and is a sister-group to the families Liwiidae and Naraoiidae.
Emucaris fava is an extinct species of soft-shelled trilobite-like arthropod of the nektaspid order from the Lower Cambrian of South Australia. It is the only species classified under the genus Emucaris.
Tariccoia is a genus of small to average size marine arthropods in the Liwiidae Family, that lived during the late Ordovician period. Fossil remains of Tariccoia were collected from Sardinia, Italy. Tariccoia looks like a large, soft agnostid trilobite. It has a headshield wider than the tailshield (pygidium), and in between them three thoracic body segments (somites). The genus is monotypic, its sole species being Tariccoia arrusensis.
Soomaspis is a genus of small to average size marine arthropods in the Liwiidae Family, that lived during the late Ordovician. Fossil remains of Soomaspis were collected from the Soom Shale Lagerstätte in Western Cape, South Africa. Soomaspis looks like a large, soft agnostid trilobite. It has a headshield wider than the tailshield (pygidium), and in between them three thoracic body segments (somites). The genus is monotypic, its sole species being Soomaspis splendida.
Pseudonaraoia is a genus of small marine arthropods within the family Naraoiidae, that lived during the late Middle Ordovician period. The only species presently known is Pseudonaraoia hammanni.
Richterops is an extinct genus of trilobite arthropods. The genus lived during the middle of the Atdabanian or the early part of the Botomian stage, which lasted from approximately 524 to 518.5 million years ago. This faunal stage was part of the second half of the Lower Cambrian. It has been found in southern Morocco. It can be recognised by the long spines of the headshield that are a smooth continuation of the frontal edge, and the enlarged spines on the 11th segment of the thorax.
Squamacula is an extinct artiopodan arthropod from the Cambrian Series 2. The type species S. clypeata was described in 1997 from the Chengjiang biota. At the time of description there were only two known specimens of S. clypeata, but now there are at least six known specimens. In 2012 a second species S. buckorum was described from the Emu Bay Shale of Australia.