Mud darter | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Order: | Perciformes |
Family: | Percidae |
Genus: | Etheostoma |
Species: | E. asprigene |
Binomial name | |
Etheostoma asprigene (S. A. Forbes, 1878) | |
Synonyms [2] | |
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The mud darter (Etheostoma asprigene) is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. [2] [3] It is endemic to the lowlands of the Mississippi River basin from Wisconsin and Minnesota south to Louisiana and East Texas. It is also found in the drainages of the Sabine and Neches Rivers of Texas and Louisiana. It can be found in slow-moving waters on riffles in rivers, as well as in creeks, swamps, lakes, and reservoirs. It mostly consumes the larvae of midges and blackflies. This species can reach a length of 7.1 cm (2.8 in), though most only reach a length of 5 cm (2.0 in).
Stephen Alfred Forbes described the mud darter in 1878. The genus name Etheostoma comes from the Greek words etheo, meaning "strain" and stoma, meaning "mouth." The species name asprigene means "rough cheek," referring to the fully scaled cheek and opercle. [2]
The mud darter has a terminal mouth with small teeth on its jaws. The back is olive or brown with 8-11 dark blotches, or saddles, across the back. The side has 9-12 dark brown irregular vertical bars and the base of the tail has three spots arranged vertically. The belly is typically a cream or light olive color. [4] They can reach a length of 7.1 cm (2.8 in), though most only reach a length of 5 cm (2.0 in).
The caudal fin is typically rounded or square in the mud darter. Its dorsal fin has two lobes, the first with 10-12 spines and the second with 10-14 rays. The mud darter does not have an adipose fin. The anal fin typically has 2 spines and 8-9 rays. [4]
The mud darter is distributed throughout the Mississippi River Basin lowlands in North America. It is found from Wisconsin and Minnesota to Louisiana and eastern Texas. It also found on the Gulf Slope in the Sabine River and Neches River drainages in Louisiana and Texas.
The mud darter is found in river mouths and overflow areas of large rivers. In these areas, it is typically found over mud covered with sand or other fine detritus. [5] They can also be found over muddy bottoms in oxbow lakes or in slow riffles of streams. Large numbers of mud darters have been seen in the finely divided tree roots along the shores of large rivers. [6] The juvenile fish are more often found in quieter areas than slow riffles.
The mud darter mainly feeds on invertebrates. The typical diet consists of midge and blackfly larvae, but some small fish were seen to have micro-crustaceans in their diet as well. [7]
Mud darters do not live over three years of age. After hatching, the fish rapidly grow to half of the total length for year 1 in ten weeks. By the end of year one, most fish averaged between 3.3 cm and 4.4 cm and by the end of years two and three, most fish averaged between 4.3 cm and 5.5 cm. Survivorship from years 1-2 was 75% for males and 81% for females, but this drastically declines, with survivorship from years 2-3 dropping to 14.5% for males and 12.1% for females. [7]
Spawning season for mud darters lasts from early March to early or mid May, with females reaching sexual maturity at one year. [7] During spawning season, males do not establish territories, but are more aggressive toward other males. During courtship, the male swims around a female with his dorsal fin erect, sometimes resting his head on her nape. The spawning site is selected by the female, while the male follows behind. The female enters the vegetation vertically and the male then positions himself over the female, curving his body into an S-shape. The pair will vibrate for a couple of seconds. The females will release between 5 and 10 eggs, which fall onto the substrate or plant. The pair will repeat this process multiple times, resting for anywhere from a few minutes to one half-hour. [7] [8]
The Bayou darter is a rare species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to western Mississippi in the United States, where it is found only in Bayou Pierre and its tributaries. Its typical habitat is fast-flowing creeks and streams, in riffles and in areas between riffles with firm gravel bottoms. It feeds on small insects and their larvae. Breeding probably takes place twice a year, in spring and late summer. The population of this fish is declining due to loss of suitable habitat. This is caused by siltation and pollution, particularly erosion caused by changes in land use. The International Union for Conservation of Nature has assessed its conservation status as being "endangered".
The harlequin darter is a species of ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae which includes the perches, ruffes and pike-perches. It is endemic to the eastern half of the United States.
The blenny darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is a poorly known species which occurs in Alabama and Tennessee where it inhabits swift riffles.
The greenside darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It inhabits swift riffles in the eastern United States and southern Ontario.
Etheostoma exile, the Iowa darter, is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is, along with about 17 other species of darters, is native to the lakes and streams of Iowa.
The Cumberland snubnose darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. This species is found in the middle Cumberland River drainage in Tennessee, Kentucky, Virginia, North Carolina, Georgia, and Alabama. It is absent in reaches above the Big South Fork, rare in North Carolina, and absent in western tributaries of the Tennessee River. While research on the ecology of E. atripinne is not extensive, what is known is they are usually found in small to medium freshwater streams in gravel riffle areas where their eggs can attach to the substrate and be left unguarded. E. atripinne can be found within a wide range of depths in its environment, leading its being classified as benthopelagic. While its global status is secure, the American Fisheries Society labels it with a status of “Special Concern”.
The teardrop darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to the eastern United States. It is only found in Kentucky and Tennessee, where it occurs in the middle to upper reaches of the Green River drainage. It inhabits small rivers and creeks and rocky pools where it feeds on the larvae of blackflies and midges, immature stages of caddisflies and mayflies, and cladocerans and copepods. This species can reach a length of 6 cm (2.4 in), though most only reach about 4.2 cm (1.7 in). This species creates nests in which the females deposit their eggs and these are guarded by the male and have been found to contain between 40 and 80 eggs. The teardrop darter was first formally described by Robert A. Kuehne and James W. Small Jr. in 1971 with the type locality given as Brush Creek, a tributary of the Green River, 2.7 miles north of Liberty, Casey County, Kentucky. The specific name honours Professor Roger W. Barbour (1919-1993) in recognition of his contribution to the knowledge of Kentucky's vertebrate fauna.
The bluntnose darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is found in eastern North America where it is found in slower moving and still waters.
The Coosa darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to the eastern United States.
The goldstripe darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to the southeastern United States where it is found in Gulf Slope streams from the Colorado River drainage in Texas to the Flint River in Georgia, the Atlantic Slope in Ocmulgee River system, Georgia, and the Mississippi embayment north as far as southeastern Missouri and western Kentucky. It is typically found in small springs, streams, and creeks with aquatic and marginal vegetation and detritus. The female spawns on multiple occasions between about mid-March and June, sticking the adhesive eggs to plants, gravel and the sides of rocks. The goldstripe darter is a common species with a wide range and numerous sub-populations, and the International Union for Conservation of Nature has assessed its conservation status as being of "least concern".
The cypress darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to fresh waters of the central and eastern United States. Its range includes drainages from the Choctawhatchee River, Florida, to the San Jacinto River, Texas, as well as the Mississippi River basin from southern Illinois and eastern Oklahoma to the Gulf of Mexico. It inhabits vegetated margins of swamps and lakes, and backwater habitats during the summer, while in winter it moves to flooded riffles and backwater bayous. It feeds mostly on small crustaceans, as well as insects and their larvae. Spawning takes place during spring and summer, with clusters of up to three eggs being laid, often stuck to the surface of dead leaves. The cypress darter has a very wide range and its population trend seems to be stable. It is a common species with numerous sub-populations, and the International Union for Conservation of Nature has assessed its conservation status as being of "least concern".
The arrow darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is found in parts of Kentucky and Tennessee, particularly in the Cumberland and Kentucky Rivers. These small fish are part of the perch family. They grow to be about 4.5 inches in length. Their opercle, breast, and cheek are all unscaled. They have a vertical black bar at the base of their caudal fin. Six to nine vertical banded bars run down the side of the fish. The dorsal side of the fish is olive green in color, and the ventral side is a yellow to white pattern. The first dorsal fin has a black base, followed by green color, and then a reddish-orange end. The second dorsal is clear with two rows of laterally lined orange spots. The caudal fin is clear and has two-three vertical lines of orange spots. The anal fin has a green base and then clear at the end. The breeding male has bright orange-red spots and breeding tubercles present on the scales.
The orangethroat darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to the central and eastern United States where it is native to parts of the Mississippi River Basin and Lake Erie Basin. Its typical habitat includes shallow gravel riffles in cooler streams and rocky runs and pools in headwaters, creeks, and small rivers, with sand, gravel, rubble, or rock substrates. It forages on the bottom for the aquatic larvae of midges, blackfly, mayfly and caddisfly, as well as isopods and amphipods. Spawning takes place in spring, the selected sites often being the upper stretches of riffles with sandy and gravelly bottoms interspersed with larger cobble. Reproductive success is high in this species. No particular threats have been identified, and the International Union for Conservation of Nature has assessed its conservation status as being of "least concern".
The Tippecanoe darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to the eastern United States.
The wounded darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to the eastern United States. Its range includes the upper Tennessee River drainage, western Virginia, western North Carolina, and eastern Tennessee. Its typical habitat is among boulders or coarse rubble and cobble, often with overhanging ledges, in medium to large slow-moving rivers. It feeds on small insect larvae, especially midge larvae. Spawning occurs when the water warms up in late spring. Females deposit clutches of adhesive eggs on the underside of rock ledges or slabs, and the male guards the nest. The population trend of this fish seems to be decreasing slowly but it is a relatively common species with numerous sub-populations, and the International Union for Conservation of Nature has assessed its conservation status as being of "least concern". The greatest threat comes from impoundment, and a management plan is in place, including captive breeding.
Etheostoma zonale, the banded darter, is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to the eastern United States. It is mainly found in the Mississippi Basin, ranging from the Verdigris River in Kansas eastward to the Allegheny River in New York, and from the Minnesota River in Minnesota southward to the Ouachita River in Arkansas and the Tennessee River in Alabama. Its typical habitat in small and medium-sized rivers is riffles over cobble or gravel, rock slabs, and small boulders. It feeds on the riverbed on small insect larvae and is itself eaten by birds and larger fish. Males become more colorful and become territorial before spawning which takes place in spring. The females attach the eggs to waterweed. The population trend of this fish seems to be stable, it is a common species with numerous sub-populations over a wide range, no major threats have been identified and the International Union for Conservation of Nature has assessed its conservation status as being of "least concern".
Percina phoxocephala, the slenderhead darter, is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is found in North America in the central Ohio and Mississippi River basins, to northeastern South Dakota and the Lake Winnebago system in Wisconsin, and as far south as the Red River in eastern Oklahoma and northeast Texas, typically in small to medium size rivers. It is a colorful species, with an average length of 6 to 9 centimeters. Males take on a deeper hue during the breeding season. It feeds on insect larvae and other small invertebrates, and spawns between April and June. It is a common fish with a very wide range and the International Union for Conservation of Nature has classified its conservation status as being of "least concern".
Etheostoma variatum, the variegate darter, is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is native to the eastern United States where it occurs primarily in the Ohio River basin; it is present in the states of Ohio, New York, Pennsylvania, Indiana, West Virginia and Kentucky with a small population in Virginia. It is a large darter, growing to 10 cm (4 in) long, orange-red with black vertical barring on its body, and horizontal barring on its front dorsal fin. Its typical habitat is swift-flowing riffles with rubble, boulder and gravel. No major threats have been identified for this fish and the International Union for Conservation of Nature has assessed its conservation status as being of "least concern". Biology
The redfin darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is native to the south-central United States, where it occurs in Missouri, Arkansas, Kansas, and eastern Oklahoma.
The gumbo darter is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is found in the Neches, Sabine, and Calcasieu River drainages in southeastern Texas and southwestern Louisiana. They inhabit riverbanks, where there are exposed roots with accumulated vegetational debris, and sand to mixed sand and gravel substrate with very little silt. This species can reach a length of 5.4 cm (2.1 in).