Notopalaeognathae | |
---|---|
Greater rhea ( Rhea americana ) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Infraclass: | Palaeognathae |
Clade: | Notopalaeognathae Yuri et al., 2013 |
Clades | |
|
Notopalaeognathae is a clade that contains the order Rheiformes (rheas), the clade Novaeratitae (which includes the cassowaries and emus, the kiwis, and the extinct elephant birds), and the clade Dinocrypturi (comprising the tinamous and the extinct moas). [3] Notopalaeognathae was named by Yuri et al. (2013) and defined in the PhyloCode by Sangster et al. (2022) as "the least inclusive crown clade containing Rhea americana , Tinamus major , and Apteryx australis ". [4] The exact relationships of this group, including its recently extinct members, have only recently been uncovered. The two lineages endemic to New Zealand, the kiwis and the extinct moas, are not each other's closest relatives: the moas are most closely related to the Neotropical tinamous, [5] [6] [7] [8] and the kiwis are sister to the extinct elephant birds of Madagascar, with kiwis and elephant birds together sister to the cassowaries and emu of New Guinea and Australia. [7] The South American rheas are either sister to all other notopalaeognaths [9] or sister to Novaeratitae. [3] The sister group to Notopalaeognathae is Struthionidae (the ostrich family).
Moa are an extinct group of flightless birds formerly endemic to New Zealand. During the Late Pleistocene-Holocene, there were nine species. The two largest species, Dinornis robustus and Dinornis novaezelandiae, reached about 3.6 metres (12 ft) in height with neck outstretched, and weighed about 230 kilograms (510 lb) while the smallest, the bush moa, was around the size of a turkey. Estimates of the moa population when Polynesians settled New Zealand circa 1300 vary between 58,000 and approximately 2.5 million.
Ratites are a polyphyletic group consisting of all birds within the infraclass Palaeognathae that lack keels and cannot fly. They are mostly large, long-necked, and long-legged, the exception being the kiwi, which is also the only nocturnal extant ratite.
Elephant birds are extinct flightless birds belonging to the order Aepyornithiformes that were native to the island of Madagascar. They are thought to have gone extinct around AD 1000, likely as a result of human activity. Elephant birds comprised three species, one in the genus Mullerornis, and two in Aepyornis.Aepyornis maximus is possibly the largest bird to have ever lived, with their eggs being the largest known for any amniote. Elephant birds are palaeognaths, and their closest living relatives are kiwi, suggesting that ratites did not diversify by vicariance during the breakup of Gondwana but instead convergently evolved flightlessness from ancestors that dispersed more recently by flying.
Flightless birds are birds that cannot fly, as they have, through evolution, lost the ability to. There are over 60 extant species, including the well-known ratites and penguins. The smallest flightless bird is the Inaccessible Island rail. The largest flightless bird, which is also the largest living bird in general, is the common ostrich.
Palaeognathae is an infraclass of birds, called paleognaths or palaeognaths, within the class Aves of the clade Archosauria. It is one of the two extant infraclasses of birds, the other being Neognathae, both of which form Neornithes. Palaeognathae contains five extant orders consisting of four flightless lineages, termed ratites, and one flying lineage, the Neotropic tinamous. There are 47 species of tinamous, five of kiwis (Apteryx), three of cassowaries (Casuarius), one of emus (Dromaius), two of rheas (Rhea) and two of ostriches (Struthio). Recent research has indicated that paleognaths are monophyletic but the traditional taxonomic split between flightless and flighted forms is incorrect; tinamous are within the ratite radiation, meaning flightlessness arose independently multiple times via parallel evolution.
Aepyornis is an extinct genus of elephant bird formerly endemic to Madagascar. The genus had two species, the smaller A. hildebrandti and the larger A. maximus, which is possibly the largest bird ever to have lived. Its closest living relative is the New Zealand kiwi. They became extinct sometime around 1000 AD, probably as a result of human activity.
Neognathae is an infraclass of birds, called neognaths, within the class Aves of the clade Archosauria. Neognathae includes the majority of living birds; the exceptions being the tinamous and the flightless ratites, which belong instead to the sister taxon Palaeognathae. There are nearly 10,000 living species of neognaths.
The grey-legged tinamou, alternatively, the gray-legged tinamou, is a small ground-dwelling bird endemic to the neotropics. It is a rarely seen bird due to its small size and discreet appearance.
The crested moa is an extinct species of moa. It is one of the 9 known species of moa to have existed.
The South Island giant moa is an extinct species of moa in the genus Dinornis, known in Māori by the name moa nunui. It was one of the tallest-known bird species to walk the Earth, exceeded in weight only by the heavier but shorter elephant bird of Madagascar.
Aequornithes, or core water birds, are defined in the PhyloCode as "the least inclusive crown clade containing Pelecanus onocrotalus and Gavia immer".
Psittacopasseres is a taxon of birds consisting of the Passeriformes and Psittaciformes (parrots). Per Ericson and colleagues, in analysing genomic DNA, revealed a lineage comprising passerines, psittacines and Falconiformes. The group was proposed following an alignment of nuclear intron sequences by Shannon Hackett et al. in 2008. It was formally named as Psittacopasserae in a 2011 Nature Communications article by Alexander Suh and other authors working with Jürgen Schmitz's group, based on genetic analysis of the insertion of retroposons into the genomes of key avian lineages over the course of evolution during the Mesozoic Era. This clade was defined in the PhyloCode by George Sangster and colleagues in 2022 as the least inclusive crown clade containing Psittacus erithacus and Passer domesticus.
Eufalconimorphae is a proposed clade of birds, consisting of passerines, parrots, falcons, caracaras, and forest falcons. It has whole-genome DNA support. This clade was defined in the PhyloCode by George Sangster and colleagues in 2022 as "the least inclusive crown clade containing Falco subbuteo and Passer domesticus". Eufalconimorphae birds are characterized by their strong and hooked beaks, sharp talons, and powerful wings. They have excellent eyesight, which allows them to spot their prey from great distances. The Eufalconimorphae is noted to produce aerodynamic force during the upstroke of flight to help create a vertical flight pattern.
Australaves is a clade of birds, defined in 2012, consisting of the Eufalconimorphae as well as the Cariamiformes. They appear to be the sister group of Afroaves. This clade was defined in the PhyloCode by George Sangster and colleagues in 2022 as "the least inclusive crown clade containing Cariama cristata and Passer domesticus".
Proapteryx micromeros is an extinct kiwi known from the 16–19 million-year-old early Miocene sediments of the St Bathans Fauna of Otago, New Zealand.
Eurypygimorphae or Phaethontimorphae is a clade of birds that contains the orders Phaethontiformes (tropicbirds) and Eurypygiformes recovered by genome analysis. The relationship was first identified in 2013 based on their nuclear genes. This group was defined in the PhyloCode by George Sangster and colleagues in 2022 as "the least inclusive crown clade containing Phaethon aethereus, Eurypyga helias, and Rhynochetos jubatus". Historically these birds were placed at different parts of the tree, with tropicbirds in Pelecaniformes and the kagu and sunbittern in Gruiformes. Some genetic analyses have placed the eurypygimorph taxa in the controversial and obsolete clade Metaves, with uncertain placement within that group. More recent molecular studies support their grouping together in Eurypygimorphae, which is usually recovered as the sister taxon to Aequornithes within Ardeae.
Otidimorphae is a clade of birds that contains the orders Cuculiformes (cuckoos), Musophagiformes (turacos), and Otidiformes (bustards) identified in 2014 by genome analysis. George Sangster and colleagues in 2022 named the clade uniting turacos and bustards as Musophagotides, defining it in the PhyloCode as "the least inclusive crown clade containing Otis tarda and Musophaga violacea, but not Grus grus or Mesitornis variegatus".
Phaethoquornithes is a clade of birds that contains Eurypygimorphae and Aequornithes, which was first recovered by genome analysis in 2014. Members of Eurypygimorphae were originally classified in the obsolete group Metaves, and Aequornithes were classified as the sister taxon to Musophagiformes or Gruiformes.
Columbimorphae is a clade/superorder discovered by genome analysis that includes birds of the orders Columbiformes, Pterocliformes (sandgrouse), and Mesitornithiformes (mesites). This group was defined in the PhyloCode by George Sangster and colleagues in 2022 as "the least inclusive crown clade containing Columba oenas, Mesitornis variegatus, and Pterocles alchata". Previous analyses had also recovered this grouping, although the exact relationships differed. Some studies indicated a sister relationship between sandgrouse and pigeons while other studies favored a sister grouping of mesites and sandgrouse instead. This sister relationship of the sandgrouses and mesites was named by George Sangster and colleagues in 2022 as the clade Pteroclimesites and defined in the PhyloCode as "the least inclusive crown clade containing Mesitornis variegatus and Pterocles alchata".
Novaeratitae is a proposed clade that was originally defined to contain the recent common ancestors of the orders Casuariiformes and Apterygiformes (kiwis). This clade was named by Yuri et al. (2013) and phylogenetically defined in the PhyloCode by Sangster et al. (2022) as "the least inclusive crown clade containing Apteryx australis and Casuarius casuarius". Recently it has been determined that the elephant birds of the extinct order Aepyornithiformes were the closest relatives of the kiwis, and therefore are part of this group. The implication is that ratites had lost flight independently in each group, as the elephant birds are the only novaeratites found outside Oceania. This clade has been contested by other studies, which find the relationships between the four main clades of non-ostrich palaeognaths to be an unresolved polytomy, with only slightly more genetic support for Novaeritiae over alternative proposals.