Palaeotis Temporal range: Eocene, | |
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Partial fossil specimen, Geisel valley museum | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Infraclass: | Palaeognathae |
Order: | Struthioniformes |
Family: | † Palaeotidae |
Genus: | † Palaeotis Lambrecht, 1928 |
Type species | |
†Palaeotis weigelti Lambrecht, 1928 | |
Synonyms [1] [2] | |
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Palaeotis is a genus of paleognath birds from the middle Eocene epoch of central Europe. One species is known, Palaeotis weigelti. The holotype specimen is a fossil tarsometatarsus and phalanx. Lambrect (1928) described it as an extinct bustard (genus Otis), and gave it its consequent name (Palaeotis means ancient bustard). [3] After a suggestion by Storrs L. Olson, a review of the type specimen and the referral of several other fossils by Houde and Haubold (1987) concluded that Palaeotis is a palaeognath and assigned it to the same order as ostriches; the Struthioniformes. [4] In 2021, it was considered a member of the family Paleotididae alongside Galligeranoides from the Early Eocene of France, which were found to be basal members of the Struthioniformes. [5]
In the 1930s a nearly complete fossil with catalog number GM 4362 was assigned to Palaeotis, probably by Lambrecht. Houde and Haubold found three additional specimens in the collection of the Geiseltalmuseum, Martin-Luther University, Halle/S., Germany. One of those three is the holotype specimen of Paleogrus geiseltalensis (=Ornithocnemus geiseltalensis, Lambrecht 1935). Houde and Haubold also requested permission to prepare a fossil cataloged as HLMD Me 7530 at the Hesseches Landesmuseum in Darmstadt, Germany. HLMD Me 7530 was collected from the famous Messel shales. When it was prepared, the two ornithologists assigned it to Palaeotis as well.
Other scientists are less convinced that Palaeotis is a struthioniform, placing it instead as a more basal ratite. It may be related to the mysterious Remiornis , a putative ratite known from the Eocene of France. Various other ratite remains also occur in the European Paleogene and early Miocene, and these may represent various independent lineages, leading to further confusion. [6] [7]
Palaeotis is suggested to be sexually dimorphic, as some specimens being consistently smaller than others. While the forelimb is incomplete, the spindley humerus indicates that it had relatively large wings unlike modern ostriches and rheas. The beak was slender, more similar to that of lithornithids, and probably had similar carnivorous habits. [8] [9] Paleotis is considered to be flightless. [10]
Struthioniformes is an order of birds with only a single extant family, Struthionidae, containing the ostriches. Several other extinct families are known, spanning across the Northern Hemisphere, from the Early Eocene to the early Pliocene, including a variety of flightless forms like the Paleotidae, Geranoididae, Eogruidae and Ergilornithidae, the latter two thought to be closely related to Struthionidae.
Ratites are a polyphyletic group consisting of all birds within the infraclass Palaeognathae that lack keels and cannot fly. They are mostly large, long-necked, and long-legged, the exception being the kiwi, which is also the only nocturnal extant ratite.
The Gruiformes are an order containing a considerable number of living and extinct bird families, with a widespread geographical diversity. Gruiform means "crane-like".
Struthionidae is a family of flightless birds, containing the extant ostriches and their extinct relatives. The two extant species of ostrich are the common ostrich and Somali ostrich, both in the genus Struthio, which also contains several species known from Holocene fossils such as the Asian ostrich. The common ostrich is the more widespread of the two living species, and is the largest living bird species. The extinct genus Pachystruthio from the Late Pliocene-Early Pleistocene of Eurasia is one of the largest birds ever.
Rheidae is a family of flightless ratite birds which first appeared in the Paleocene. It is today represented by the sole living genus Rhea, but also contains several extinct genera.
The mousebirds are birds in the order Coliiformes. They are the sister group to the clade Cavitaves, which includes the Leptosomiformes, Trogoniformes (trogons), Bucerotiformes, Piciformes and Coraciformes. This group is now confined to sub-Saharan Africa, and it is the only bird order confined entirely to that continent, with the possible exception of turacos which are considered by some as the distinct order Musophagiformes, and the cuckoo roller, which is the only member of the order Leptosomiformes, and which is found in Madagascar but not mainland Africa. Mousebirds had a wider range in the Paleogene, with a widespread distribution in Europe and North America during the Paleocene.
Ostriches are large flightless birds. Two living species are recognised, the common ostrich, native to large areas of sub-Saharan Africa, and the Somali ostrich, native to the Horn of Africa.
Palaeognathae is an infraclass of birds, called paleognaths or palaeognaths, within the class Aves of the clade Archosauria. It is one of the two extant infraclasses of birds, the other being Neognathae, both of which form Neornithes. Palaeognathae contains five extant orders consisting of four flightless lineages, termed ratites, and one flying lineage, the Neotropic tinamous. There are 47 species of tinamous, five of kiwis (Apteryx), three of cassowaries (Casuarius), one of emus (Dromaius), two of rheas (Rhea) and two of ostriches (Struthio). Recent research has indicated that paleognaths are monophyletic but the traditional taxonomic split between flightless and flighted forms is incorrect; tinamous are within the ratite radiation, meaning flightlessness arose independently multiple times via parallel evolution.
Strigogyps is an extinct genus of prehistoric bird from the Middle Eocene to Early Oligocene of France and Germany. It was probably around the size of a large chicken or a guan, weighing not quite 1 kilogram (2.2 lb). Apparently, as indicated by the ratio of lengths of wing to leg bones, S. sapea was flightless. Its legs were not adapted to running, so it seems to have had a walking lifestyle similar to trumpeters. Unlike other Cariamiformes which appear to have been mostly carnivorous, the specimens of one species, Strigogyps sapea, suggest a facultatively herbivorous or omnivorous diet.
Diogenornis is an extinct genus of ratites, that lived from the Middle Paleocene to the Early Eocene. It was described in 1983 by Brazilian scientist Herculano Marcos Ferraz de Alvarenga based on fossils found in the Itaboraí Formation in southeastern Brazil. The type species is D. fragilis. It grew to about two thirds the size of the modern greater rhea, at about 90 centimetres (3.0 ft) of height.
Dasornis is a genus of prehistoric pseudotooth birds. These were probably close relatives of either pelicans and storks or waterfowl; they are placed in the order Odontopterygiformes to account for this uncertainty.
Macrodontopteryx is a genus of the prehistoric pseudotooth birds of somewhat doubtful validity. These animals were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Lithornis is a genus of extinct paleognathous birds. Although Lithornis was able to fly well, their closest relatives are the extant tinamous and ratites.
Lithornithidae is an extinct, possibly paraphyletic group of early paleognath birds. They are known from fossils dating to the Upper Paleocene through the Middle Eocene of North America and Europe, with possible Late Cretaceous representatives. All are extinct today; the youngest specimen is the currently unnamed SGPIMH MEV1 specimen from the mid-Eocene Messel Pit site.
Eremopezus is a prehistoric bird genus, possibly a palaeognath. It is known only from the fossil remains of a single species, the huge and presumably flightless Eremopezus eocaenus. This was found in Upper Eocene Jebel Qatrani Formation deposits around the Qasr el Sagha escarpment, north of the Birket Qarun lake near Faiyum in Egypt. The rocks its fossils occur in were deposited in the Priabonian, with the oldest dating back to about 36 million years ago (Ma) and the youngest not less than about 33 Ma.
Pangalliformes is the scientific name of a provisional clade of birds within the group Galloanserae. It is defined as all birds more closely related to chickens than to ducks, and includes all modern chickens, turkeys, pheasants, and megapodes, as well as extinct species that do not fall within the crown group Galliformes.
Cariamiformes is an order of primarily flightless birds that has existed for over 50 million years. The group includes the family Cariamidae (seriemas) and the extinct families such as Phorusrhacidae, Bathornithidae, Idiornithidae and Ameghinornithidae. Extant members (seriemas) are only known from South America, but fossils of many extinct taxa are also found in other continents including Europe and North America. Though traditionally considered a suborder within Gruiformes, both morphological and genetic studies show that it belongs to a separate group of birds, Australaves, whose other living members are Falconidae, Psittaciformes and Passeriformes.
Eleutherornis cotei is an extinct flightless predatory cariamiform bird which lived during the Middle Eocene of France and Switzerland. Since the early 20th century, researchers have initially described the fossils of Eleutherornis as separate taxa, some remains as a species of Gastornis and others as an ancient ratite related to modern ostriches. However, subsequent analyses have questioned the original interpretations, and a thorough reexamination in 2013 indicated that all of these described remains represent the same species.
Eogruidae is a family of large, flightless birds that inhabited Asia from the Eocene to Pliocene epochs. Related to modern ostriches, it was formerly thought to be related to cranes, limpkins and trumpeters and that the similarities with ostriches were due to similar speciations to cursoriality, with both groups showing reduced numbers of toes to two in some taxa. It has been suggested that competition from true ostriches has caused the extinction of these birds, though this has never been formally tested and several ostrich taxa do occur in the late Cenozoic of Asia and some species do occur in areas where ostrich fossils have also been found. It has been suggested that the family is paraphyletic, with Ergilornithidae more closely related to modern ostriches than to Eogrus or Sonogrus.
Geranoididae is a clade of extinct birds from the early to late Eocene and possibly early Oligocene of North America and Europe. These were mid-sized, long-legged flightless birds. Recent research shows that these birds may actually be palaeognaths related to ostriches.