Eogruidae Temporal range: | |
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Detailed depiction of Ergilornis in its habitat | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Infraclass: | Palaeognathae |
Order: | Struthioniformes |
Family: | † Eogruidae Wetmore, 1934 |
Genera | |
Eogruidae (also spelled Eogruiidae in some publications) is a family of large, flightless birds that inhabited Asia from the Eocene to Pliocene epochs. Related to modern ostriches, it was formerly thought to be related to cranes, limpkins and trumpeters and that the similarities with ostriches were due to similar speciations to cursoriality, with both groups showing reduced numbers of toes to two in some taxa. [1] [2] [3] It has been suggested that competition from true ostriches has caused the extinction of these birds, [2] [3] though this has never been formally tested and several ostrich taxa do occur in the late Cenozoic of Asia [4] and some species do occur in areas where ostrich fossils have also been found. [1] It has been suggested that the family is paraphyletic, with Ergilornithidae more closely related to modern ostriches than to Eogrus or Sonogrus . [5]
Most eogruids are known from rather sparse remains, mostly the tarso-metatarsals and toes. The former are generally slender with a distinct crest along the lateral side of the plantar surface. The trochlea for the second toe shows a progressive reduction along the various taxa, culminating in its utter absence in Amphipelargus. [4] [3]
Other skeletal remains are rare. Wing elements are known in a few taxa; these are generally highly reduced, suggesting that these birds were flightless. The exception appears to be Eogrus itself, which does not show much phalange reduction and hypothetically could still fly, [4] though the rest of the postcranial skeleton is already specialised for cursoriality. [2] [3]
Eogruids were formerly considered to be Gruiformes within the crane-limpkin-trumpeter line, Gruoidea, the exception being Olson 1985 which declared them to be stem-ostriches. [4] A study in 2021 based on newly described remains found that eogruids and the related Ergilornithidae are indeed members of Struthoniformes. [6]
More derived taxa such as Ergilornis and Amphipelargus were sometimes classified as a separate family, Ergilornithidae, but they are now generally accepted to be a subfamily within Eogruidae. [1] Eogruids are rather similar to Geranoididae, a clade of similar flightless gruiforms from North America and Europe, and both groups have occasionally been classified as sister taxa. Recently geranoidids have been recovered as basal to the rest of Gruoidea, however, while eogruids are sister-taxa to cranes. [7]
There is some contention in regards to ergilornithine genera, particularly Amphipelargus and Urmiornis, which have assimilated each other's species from study to study. The most recent consensus is that nearly all Neogene eogruid remains belong to Urmiornis, while a few western Eurasian taxa can be referred to as Amphipelargus. [1]
Struthioniformes is an order of birds with only a single extant family, Struthionidae, containing the ostriches. Several other extinct families are known, spanning across the Northern Hemisphere, from the Early Eocene to the early Pliocene, including a variety of flightless forms like the Paleotididae, Geranoididae, Eogruidae and Ergilornithidae, the latter two thought to be closely related to Struthionidae.
The Gruiformes are an order containing a considerable number of living and extinct bird families, with a widespread geographical diversity. Gruiform means "crane-like".
Rails are a large, cosmopolitan family of small- to medium-sized terrestrial and/or semi-amphibious birds. The family exhibits considerable diversity in its forms, and includes such ubiquitous species as the crakes, coots, and gallinule; other rail species are extremely rare or endangered. Many are associated with wetland habitats, some being semi-aquatic like waterfowl, but many more are wading birds or shorebirds. The ideal rail habitats are marsh areas, including rice paddies, and flooded fields or open forest. They are especially fond of dense vegetation for nesting. The rail family is found in every terrestrial habitat with the exception of dry desert, polar or freezing regions, and alpine areas. Members of Rallidae occur on every continent except Antarctica. Numerous unique island species are known.
Struthionidae is a family of flightless birds, containing the extant ostriches and their extinct relatives. The two extant species of ostrich are the common ostrich and Somali ostrich, both in the genus Struthio, which also contains several species known from Holocene fossils such as the Asian ostrich. The common ostrich is the more widespread of the two living species, and is the largest living bird species. The extinct genus Pachystruthio from the Late Pliocene-Early Pleistocene of Eurasia is one of the largest birds ever.
The mousebirds are birds in the order Coliiformes. They are the sister group to the clade Cavitaves, which includes the Leptosomiformes, Trogoniformes (trogons), Bucerotiformes, Piciformes and Coraciformes. This group is now confined to sub-Saharan Africa, and it is the only bird order confined entirely to that continent, with the possible exception of turacos which are considered by some as the distinct order Musophagiformes, and the cuckoo roller, which is the only member of the order Leptosomiformes, and which is found in Madagascar but not mainland Africa. Mousebirds had a wider range in the Paleogene, with a widespread distribution in Europe and North America during the Paleocene.
Ostriches are large flightless birds. Two living species are recognised, the common ostrich, native to large areas of sub-Saharan Africa, and the Somali ostrich, native to the Horn of Africa.
Phorusrhacids, colloquially known as terror birds, are an extinct family of large carnivorous, mostly flightless birds that were among the largest apex predators in South America during the Cenozoic era. Their definitive fossil records range from the Middle Eocene to the Late Pleistocene around 43 to 0.1 million years ago, though some specimens suggest that they were present since the Early Eocene.
Palaeognathae is an infraclass of birds, called paleognaths or palaeognaths, within the class Aves of the clade Archosauria. It is one of the two extant infraclasses of birds, the other being Neognathae, both of which form Neornithes. Palaeognathae contains five extant branches of flightless lineages, termed ratites, and one flying lineage, the Neotropic tinamous. There are 47 species of tinamous, five of kiwis (Apteryx), three of cassowaries (Casuarius), one of emus (Dromaius), two of rheas (Rhea) and two of ostriches (Struthio). Recent research has indicated that paleognaths are monophyletic but the traditional taxonomic split between flightless and flighted forms is incorrect; tinamous are within the ratite radiation, meaning flightlessness arose independently multiple times via parallel evolution.
Strigogyps is an extinct genus of prehistoric bird from the Middle Eocene to Early Oligocene of France and Germany. It was probably around the size of a large chicken or a guan, weighing not quite 1 kilogram (2.2 lb). Apparently, as indicated by the ratio of lengths of wing to leg bones, S. sapea was flightless. Its legs were not adapted to running, so it seems to have had a walking lifestyle similar to trumpeters. Unlike other Cariamiformes which appear to have been mostly carnivorous, the specimens of one species, Strigogyps sapea, suggest a facultatively herbivorous or omnivorous diet.
Palaeotis is a genus of paleognath birds from the middle Eocene epoch of central Europe. One species is known, Palaeotis weigelti. The holotype specimen is a fossil tarsometatarsus and phalanx. Lambrect (1928) described it as an extinct bustard, and gave it its consequent name. After a suggestion by Storrs L. Olson, a review of the type specimen and the referral of several other fossils by Houde and Haubold (1987) concluded that Palaeotis is a palaeognath and assigned it to the same order as ostriches; the Struthioniformes. In 2021, it was considered a member of the family Paleotididae alongside Galligeranoides from the Early Eocene of France, which were found to be basal members of the Struthioniformes.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 1976.
Macrodontopteryx is a genus of the prehistoric pseudotooth birds of somewhat doubtful validity. These animals were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Lithornithidae is an extinct, possibly paraphyletic group of early paleognath birds. They are known from fossils dating to the Upper Paleocene through the Middle Eocene of North America and Europe, with possible Late Cretaceous representatives. All are extinct today; the youngest specimen is the currently unnamed SGPIMH MEV1 specimen from the mid-Eocene Messel Pit site.
Eremopezus is a prehistoric bird genus, possibly a palaeognath. It is known only from the fossil remains of a single species, the huge and presumably flightless Eremopezus eocaenus. This was found in Upper Eocene Jebel Qatrani Formation deposits around the Qasr el Sagha escarpment, north of the Birket Qarun lake near Faiyum in Egypt. The rocks its fossils occur in were deposited in the Priabonian, with the oldest dating back to about 36 million years ago (Ma) and the youngest not less than about 33 Ma.
Gruimorphae is a clade of birds that contains the orders Charadriiformes and Gruiformes identified by molecular analysis. This grouping has had historical support, as various charadriiform families such as the families Pedionomidae and Turnicidae were classified as gruiforms. It may also have support from the fossil record since the discovery of Nahmavis from the Early Eocene of North America.
Bathornithidae is an extinct family of birds from the Eocene to Miocene of North America. Part of Cariamiformes, they are related to the still extant seriemas and the extinct Phorusrhacidae. They were likely similar in habits, being terrestrial, long-legged predators, some of which attained massive sizes.
Eutreptornis is a genus of extinct possible cariamiforme bird from the Late Eocene of Utah. It is traditionally considered to be a bathornithid, though a combination of the relative incompleteness of the material alongside some differences from other bathornithids have raised some suspicions about this affiliation.
Bathornis is an extinct lineage of birds related to modern day seriemas, that lived in North America about 37–20 million years ago. Like the closely related and also extinct phorusrhacids, it was a flightless predator, occupying predatory niches in environments classically considered to be dominated by mammals. It was a highly diverse and successful genus, spanning a large number of species that occurred from the Priabonian Eocene to the Burdigalian Miocene epochs.
Geranoididae is a clade of extinct birds from the early to late Eocene and possibly early Oligocene of North America and Europe. These were mid-sized, long-legged flightless birds. Recent research shows that these birds may actually be palaeognaths related to ostriches.
Messelornithidae is an extinct clade of gruiform birds, closely related to modern rails. The fossil record are from the Paleocene to the early Oligocene of Europe and North America.