Chitons are marine molluscs of varying size in the class Polyplacophora, formerly known as Amphineura. About 940 extant and 430 fossil species are recognized.
Monoplacophora, meaning "bearing one plate", is a polyphyletic superclass of molluscs with a cap-like shell inhabiting deep sea environments. Extant representatives were not recognized as such until 1952; previously they were known only from the fossil record, and were thought to have become extinct 375 million years ago.
Wiwaxia is a genus of soft-bodied animals that were covered in carbonaceous scales and spines that protected it from predators. Wiwaxia fossils – mainly isolated scales, but sometimes complete, articulated fossils – are known from early Cambrian and middle Cambrian fossil deposits across the globe. The living animal would have measured up to 5 cm (2 inch) when fully grown, although a range of juvenile specimens are known, the smallest being 2 millimetres (0.079 in) long.
The tusk shells or tooth shells, technically the Scaphopoda, are members of a class of shelled marine mollusc with worldwide distribution, and are the only class of exclusively infaunal marine molluscs. Shells of species within this class range from about 0.5 to 18 cm in length. Members of the order Dentaliida tend to be larger than those of the order Gadilida.
Aplacophora is a presumably paraphyletic taxon. This is a class of small, deep-water, exclusively benthic, marine molluscs found in all oceans of the world.
The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is Halkieria, which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the small shelly fossil assemblages. The best known species is Halkieria evangelista, from the North Greenland Sirius Passet Lagerstätte, in which complete specimens were collected on an expedition in 1989. The fossils were described by Simon Conway Morris and John Peel in a short paper in 1990 in the journal Nature. Later a more thorough description was undertaken in 1995 in the journal Philosophical Transactions of the Royal Society of London and wider evolutionary implications were posed.
The evolution of the molluscs is the way in which the Mollusca, one of the largest groups of invertebrate animals, evolved. This phylum includes gastropods, bivalves, scaphopods, cephalopods, and several other groups. The fossil record of mollusks is relatively complete, and they are well represented in most fossil-bearing marine strata. Very early organisms which have dubiously been compared to molluscs include Kimberella and Odontogriphus.
Odontogriphus is a genus of soft-bodied animals known from middle Cambrian Lagerstätte. Reaching as much as 12.5 centimetres (4.9 in) in length, Odontogriphus is a flat, oval bilaterian which apparently had a single muscular foot and a "shell" on its back that was moderately rigid but of a material unsuited to fossilization.
Halwaxiida or halwaxiids is a proposed clade equivalent to the older orders Sachitida He 1980 and Thambetolepidea Jell 1981, loosely uniting scale-bearing Cambrian animals, which may lie in the stem group to molluscs or lophotrochozoa. Some palaeontologists question the validity of the Halwaxiida clade.
Mollusca is the second-largest phylum of invertebrate animals, after the Arthropoda; members are known as molluscs or mollusks. Around 85,000 extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied.
Matthevia is a genus of Cambrian molluscs, perhaps related to the chitons. It consists of repeated monoplacophoran-like shells; according to one hypothesis, chitons arose when these tall shells began to overlap over the generations. The tall element of the shell was retained and forms the tips of modern chiton plates. There are distinct head, 'centre', and tail valves, which occur approximately in the ratio 1:5:1 — suggesting a seven-plated configuration.
Heloplax is a genus of worm-like molluscs. Its soft parts are preserved in three dimensions in the Silurian Herefordshire Lagerstatte; its disarticulated valves are known from other Silurian deposits. It is very bizarre by modern standards; it bears serially repeated units, and has spines. It probably falls somewhere between the aplacophorans and polyplacophora; its valves were composed of aragonite
Acaenoplax is an extinct worm-shaped mollusc known from the Coalbrookdale Formation of Herefordshire, England. It lived in the Silurian period. It was a couple of centimetres long and half a centimetre wide, and comprises serially repeated units with seven or eight shells, and rings of 'spines'.
The origin of the brachiopods is uncertain; they either arose from reduction of a multi-plated tubular organism, or from the folding of a slug-like organism with a protective shell on either end. Since their Cambrian origin, the phylum rose to a Palaeozoic dominance, but dwindled during the Mesozoic.
The cephalopods have a long geological history, with the first nautiloids found in late Cambrian strata, and purported stem-group representatives present in the earliest Cambrian lagerstätten.
Hanleya is a genus of polyplacophoran molluscs known from Oligocene and Miocene fossils; it is represented today by a number of species including H. sinica Xu 1990 (China), H. brachyplax (Brazil) and H. hanleyi Bean in Thorpe, 1844 (Chile), which feeds on sponges.
Helminthochiton is an extinct genus of polyplacophoran mollusc. Helminthochiton became extinct during the Permian period.
Acutichiton is among the most primitive genera of Neoloricate chitons. Acutichiton became extinct during the Carboniferous period. Articulated specimens are known.
The Heloplacidae are a group of plated aplacophora known from Silurian deposits. Their best understood representative, Acaenoplax, can be taken as representative of the family; it is the only genus for which soft part anatomy is known.
Kulindroplax perissokomos is a Silurian mollusk, known from a single fossil from the Coalbrookdale Formation fauna of England. It lived during the Homerian Age. It is considered a basal aplacophoran. Unlike all modern aplacophorans, which are shell-less, Kulindroplax has a chiton-like shell, and it is considered a transitional fossil in the evolution of molluscs.
