Hornets are the largest of the eusocial wasps, and are similar in appearance to their close relatives yellowjackets. Some species can reach up to 5.5 cm (2.2 in) in length. They are distinguished from other vespine wasps by the relatively large top margin of the head and by the rounded segment of the abdomen just behind the waist. Worldwide, 22 species of Vespa are recognized. Most species only occur in the tropics of Asia, though the European hornet, is widely distributed throughout Europe, Russia, North America, and Northeast Asia. Wasps native to North America in the genus Dolichovespula are commonly referred to as hornets, but are actually yellowjackets.
Wasps of the cosmopolitan genus Polistes are the most familiar of the polistine wasps, and are the most common type of paper wasp in North America. Walter Ebeling coined the vernacular name "umbrella wasps" for this genus in 1975 to distinguish it from other types of paper wasp, in reference to the form of their nests. It is also the single largest genus within the family Vespidae, with over 300 recognized species and subspecies. Their innate preferences for nest-building sites leads them to commonly build nests on human habitation, where they can be very unwelcome; although generally not aggressive, they can be provoked into defending their nests. All species are predatory, and they may consume large numbers of caterpillars, in which respect they are generally considered beneficial. The European paper wasp, Polistes dominula, was introduced into the US about 1981 and has quickly spread throughout most of the country, in most cases replacing native species within a few years. This species is very commonly mistaken for a yellow jacket, as it is black, strongly marked with yellow, and quite different from the native North American species of Polistes. The cuckoo wasp, Polistes sulcifer, is an obligate social parasite, whose only host is P. dominula. Polistes annularis, whose species name is Latin for "ringed", is also known for its distinctive red body color. Polistes metricus adults malaxate their insect prey by chewing them into a pulp, sucking out and ingesting the body fluids, then feeding the rest of the morsel to their larvae. The most widely distributed South American wasp species, Polistes versicolor, is particularly common in the southeastern Brazilian states. This social wasp is commonly referred to as the yellow paper wasp due to the distinct yellow bands found on its thorax and abdomen. Polistes wasps can be identified by their characteristic flight; their long legs dangle below their bodies, which are also more slender than a yellow jacket.
The Oriental hornet is a social insect species of the family Vespidae. It can be found in Southwest Asia, Northeast Africa, the island of Madagascar, the Middle East, Central Asia, and parts of Southern Europe. Oriental hornets have also been found in a few isolated locations such as Mexico and Chile due to human introduction. The Oriental hornet lives in seasonal colonies consisting of caste system dominated by a queen. The hornet builds its nests underground and communicates using sound vibrations. The hornet has a yellow stripe on its cuticle (exoskeleton), which can absorb sunlight to generate a small electrical potential, and this might help supply energy for digging. The adult hornet eats nectar and fruits and scavenges for insects and animal proteins to feed to its young. Because they are scavengers, the hornets may also serve as a transmitter of disease following consumption of infected plants. The hornets are a primary pest to honey bees, attacking bee colonies to obtain honey and animal proteins. The sting of an Oriental hornet can be quite painful to humans and some humans are allergic to stings.
Vespula vulgaris, known as the common wasp, is a species found in regions that include the United Kingdom, Germany, India, China, New Zealand and Australia. It is sometimes known in English as the European wasp, but the same name is used for the species Vespula germanica or German wasp. Another name for Vespula vulgaris is the common yellow-jacket. In 2010, the ostensible Vespula vulgaris wasps in North America were found to be a different species, Vespula alascensis.
Polistes gallicus is a species of paper wasp found in various parts of Europe, excluding England, Denmark, and Scandinavia, from warmer climates to cooler regions north of the Alps. Nests of these social insects are created in these various conditions. The Polistes species use an oral secretion to construct their nests, which consist of a combination of saliva and chewed plant fibers. This structural mixture physically protects the nest from various harsh elements and from weathering over time.
Polistes humilis, known as the common paper wasp, is a species of wasp in the Vespidae family that is found throughout Australia and which has been introduced to northern New Zealand. These paper wasps can be identified by their long thin legs and banded yellow and black coloring. They have been known to re-utilize old nests. While the species does not exhibit morphological class differences, there are distinct behavioral differences between queens and workers. In addition, the species is eusocial and benefits from relatedness between individuals. They are known for delivering a painful sting, especially when their nest is disturbed, a behavior that has been developed as a nest defense mechanism. While wasps are often viewed negatively, they play an important pollination role for many plants.
Ropalidia marginata is an Old World species of paper wasp. It is primitively eusocial, not showing the same bias in brood care seen in other social insects with greater asymmetry in relatedness. The species employees a variety of colony founding strategies, sometimes with single founders and sometimes in groups of variable number. The queen does not use physical dominance to control workers; there is evidence of pheromones being used to suppress other female workers from overtaking queenship.
Polistes chinensis is a polistine vespid wasp in the cosmopolitan genus Polistes, and is commonly known as the Asian, Chinese or Japanese paper wasp. It is found in East Asia, in particular China and Japan. The subspecies P. chinensis antennalis is an invasive species in New Zealand, having arrived in 1979.
Polistes annularis is a species of paper wasp found throughout the eastern half of the United States. This species of red paper wasp is known for its large size and its red-and-black coloration and is variably referred to as a ringed paper wasp or jack Spaniard wasp. It builds its nest under overhangs near bodies of water that minimize the amount of sunlight penetration. It clusters its nests together in large aggregations, and consumes nectar and other insects. Its principal predator is the ant, although birds are also known to prey on it. Unlike other wasps, P. annularis is relatively robust in winter conditions, and has also been observed to store honey in advance of hibernation. This species has also been used as a model species to demonstrate the ability to use microsatellite markers in maternity assignment of social insects.
Polistes metricus is a wasp native to North America. In the United States, it ranges throughout the southern Midwest, the South, and as far northeast as New York, but has recently been spotted in southwest Ontario. A single female specimen has also been reported from Dryden, Maine. Polistes metricus is dark colored, with yellow tarsi and black tibia. Nests of Polistes metricus can be found attached to the sides of buildings, trees, and shrubbery.
Polistes fuscatus, whose common name is the dark or northern paper wasp, is widely found in eastern North America, from southern Canada through the southern United States. It often nests around human development. However, it greatly prefers areas in which wood is readily available for use as nest material, therefore they are also found near and in woodlands and savannas. P. fuscatus is a social wasp that is part of a complex society based around a single dominant foundress along with other cofoundresses and a dominance hierarchy.
Dolichovespula arenaria, also known as the common aerial yellowjacket, sandhills hornet, and common yellow hornet, is a species of wasp within the genus Dolichovespula widely distributed in the North American continent.
Polistes snelleni, the Japanese paper wasp, is a common social wasp species in central and northern Japan. P. snelleni is also found in northern China, Korea, and the Russian Far East. Due to the different climates in these regions, P. snelleni is able to adapt to different temperatures and climatic conditions. P. snelleni is typically found in hilly or submontane areas, so they are classified in the semi-highland category.
Polistes canadensis is a species of red paper wasp found in the Neotropical realm. It is a primitively eusocial wasp as a member of the subfamily Polistinae. A largely predatory species, it hunts for caterpillar meat to supply its colony, often supplementing its developing larvae with nectar. The most widely distributed American species of the genus Polistes, it colonises multiple combs, which it rears year-round.
Agelaia pallipes is a species of social paper wasp found from Costa Rica to Argentina and Paraguay. A. pallipes is ground-nesting and is one of the most aggressive wasps in South America. This species is a predator of other insects, including flies, moths, and ground crickets, as well as baby birds.
Polistes japonicus is a eusocial paper wasp found in Japan. It was first described by Henri Louis Frédéric de Saussure in 1858. It is closely related to Polistes formosanus. This species lives in small colonies with few workers and a foundress queen. Nests of these wasps are sometimes used as a traditional medicine in Korea, China, and Japan.
Polistes versicolor, also known as the variegated paper wasp or yellow paper wasp, is a subtropical social wasp within Polistes, the most common genus of paper wasp. The most widely distributed South American wasp species, P. versicolor is particularly common in the Southeastern Brazilian states. This social wasp is commonly referred to as the yellow paper wasp due to the distinct yellow bands found on its thorax and abdomen. The P. versicolor nest, made of chewed vegetable fiber, is typically a single, uncovered comb attached to the substratum by a single petiole. The yellow wasp is frequently found in urban areas. New nests and colonies are usually founded by an association of females, sometimes in human buildings. The P. versicolor colony cycle broadly ranges from 3 to 10 months, although there appears to be no relationship between the colony's development and the season of the year. While yellow paper wasps do have clear annual colony cycles, many young queens have the opportunity to hibernate during the winter, forming optional winter aggregations. Dominance hierarchies within these aggregations are characterized by physical aggression of the dominant female(s) towards the associated females, who tend to be sisters. Wagging movements are also often used as a form of communication within the colony. The yellow paper wasp is generally predatory, capturing a wide range of insects, although it often feeds on pollen and nectar as well. Therefore, P. versicolor can be useful as a pollinator or as effective pest control.
Polistes dorsalis is a species of social wasps that can be found throughout various parts of North America. It is classified under the Vespidae within the genus of Polistes. Male Polistes dorsalis wasps can be distinguished from other Polistes species by their distinctly prominent median tubercle of sternum 7. Both sexes can also be recognized due to their v- shaped yellow markings on their head. They are distributed widely across North America and can be found in sheltered nests, typically closer to the ground. These wasps live in a dominance hierarchy in which the queen's role differs from that of ordinary workers. When threatened, these wasps can deliver moderately painful stings. Their venom might also be of human interest for their antimicrobial uses.
Polistes erythrocephalus is a species of paper wasp in the subfamily Polistinae of family Vespidae found in Central and South America. P. erythrocephalus is a eusocial wasp, meaning that it possesses both reproductive and non-reproductive castes. The cooperation between the two castes to raise young demonstrates the altruistic nature of these wasps. P. erythrocephalus exhibits a four-stage colony cycle, as do many other Polistes wasps. This species generally feeds on larvae, occasionally their own, and is preyed upon by species such as army ants.
Mischocyttarus mexicanus cubicola is a neotropical paper wasp found in the New World. It is a social wasp that demonstrates two different types of nesting strategies, depending upon context. This context-dependent trait makes Mischocyttarus mexicanus cubicola a good model to study social biology within social wasps. In detail, this trait allows for the females of this species to form nests both individually and as co-founders with other females within the same colony. This subspecies is also known to exhibit cannibalism, with M. m. cubicola queens feeding on their own larvae for nourishment when unaided by workers.
