Productidae Temporal range: | |
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Fossil of Productus crawfordsvillensis from the Monte Cristo Group | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Brachiopoda |
Class: | † Strophomenata |
Order: | † Productida |
Superfamily: | † Productoidea |
Family: | † Productidae Gray, 1840 |
Subgroups | |
Productidae is an extinct family of brachiopods which lived from the Upper Devonian to Upper Permian periods in marine environments. [1] It is the most diversified family in the suborder Productidina, with some 100 genera. [2]
The exact evolutionary relationships of Productidae as well as which groups belong to this family have been a matter of extensive debate throughout much of the 20th Century, primarily due to the three genera Buxtonia , Pustula and Juresania (with the debate later expanded to their families and subfamilies) shifting in position repeatedly between phylogenies and classifications. The emphasis on internal versus external characters to determine the systematics of these groups has largely been responsible for this: the original Treatise on Invertebrate Paleontology published in 1965 placed emphasis on internal characters (including the cardinal process) as diagnostic, whereas the 2000 revision primarily used external features and shell shape, resulting in differing classification of these clades. [3]
More recently, Leighton & Maples (2002) conducted multiple phylogenetic analyses which are strongly in agreement that Buxtoniinae and Juresaniinae belong in Echinoconchidae rather than Productidae, and that Productidae contains only the subfamilies Leioproductinae, Dictyoclostinae and Productinae. In addition, the genus Setigerites (historically placed in Buxtoniinae) has features of both productids and echinoconchids, potentially being related to the common ancestor of the two families. The cladogram results of their phylogenetic analyses are displayed in the cladogram below: [4]
Members of this family had a deep corpus cavity (rarely moderate or shallow) and inflated ventral corpus, with rows of spines near the hinge. Hinge teeth are absent except in the oldest genera. [1] [4] The cardinal process of productids can be split into two main types: bi-lobed or tri-lobed. [5]
Strophomenida is an extinct order of articulate brachiopods which lived from the lower Ordovician period to the mid Carboniferous period. Strophomenida is part of the extinct class Strophomenata, and was the largest known order of brachiopods, encompassing over 400 genera. Some of the largest and heaviest known brachiopod species belong to this class. Strophomenids were among the most diverse and abundant brachiopods during the Ordovician, but their diversity was strongly impacted at the Late Ordovician mass extinction. Survivors rediversified into new morphologies in the Silurian, only to be impacted once again at the Late Devonian mass extinction. However, they still survived till the mid Carboniferous.
The extinct Solenoporaceae have traditionally been interpreted as a group of red algae ancestral to the Corallinales.
Rhynchonelliformea is a major subphylum and clade of brachiopods. It is roughly equivalent to the former class Articulata, which was used previously in brachiopod taxonomy up until the 1990s. These so-called articulated brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations and separate sets of simple opening and closing muscles.
Pterodon is an extinct genus of superficially wolf-like hyainailurids from subtribe Pterodontina within polyphyletic tribe Hyainailourini in paraphyletic subfamily Hyainailourinae, that lived in late Eocene Europe.
Phosphorosaurus is an extinct genus of marine lizard belonging to the mosasaur family. Phosphorosaurus is classified within the Halisaurinae subfamily alongside the genera Pluridens, Eonatator, and Halisaurus.
Productida is an extinct order of brachiopods in the extinct class Strophomenata. Members of Productida first appeared during the Silurian. They represented the most abundant group of brachiopods during the Permian period, accounting for 45-70% of all species. The vast majority of species went extinct during the Permian-Triassic extinction event, though a handful survived into the Early Triassic. Many productids are covered in hollow tubular spines, which are characteristic of the group. A number of functions for the spines have been proposed, including as a defensive mechanism against predators.
Kutorginates (Kutorginata) are an extinct class of early rhynchonelliform ("articulate") brachiopods. The class contains only a single order, Kutorginida (kutorginides). Kutorginides were among the earliest rhynchonelliforms, restricted to the lower-middle part of the Cambrian Period.
The orthotetides (Orthotetida) are an extinct order of brachiopods in the class Strophomenata. Though not particularly diverse or abundant relative to strophomenides (Strophomenida) or productides (Productida), orthotetides were nevertheless the longest-lasting order of strophomenates, surviving from the Middle Ordovician (“Llanvirn”) up until the Late Permian. Externally, many orthotetides are difficult to distinguish from strophomenides. Most fundamental differences between the two orders are internal: orthotetides have more elaborate cardinal processes and a greater diversity of shell microstructure.
Diaphragmus is an extinct genus of brachiopod belonging to the order Productida and family Linoproductidae. Specimens have been found in Carboniferous beds in North America.
Pugnoides is an extinct genus of brachiopod belonging to the order Rhynchonellida and family Petasmariidae. Specimens have been found in Devonian to Permian beds in North America, Asia, Europe, western Australia, New Zealand,and New Zealand. The genus was particularly widespread in the Visean.
Interatheriinae is an extinct subfamily of interatheriids that consisted of notoungulates dating from the Early Eocene to the Early Pliocene. The subfamily includes the genera Archaeophylus, Argyrohyrax, Boleatherium, Brucemacfaddenia, Caenophilus, Choichephilum, Cochilius, Eopachyrucos, Federicoanaya, Interatherium, Juchuysillu, Miocochilius, Neoicochilus, Patriarchus, Proargyrohyrax, Progaleopithecus, Protypotherium, and Santiagorothia. They were small to medium sized interatheres, and when compared to the other subfamily, Notopithecinae, interatheriines are found to occupy an advanced, derived position in the family.
Echinoconchidae is an extinct family of brachiopods which lived from the Lower Carboniferous to Upper Permian periods in marine habitats. Currently, four subfamilies are assigned to it, though the evolutionary relationships between them and the family Productidae have been heavily debated for the better part of the 20th Century.
Echinoconchinae is an extinct subfamily of brachiopods which lived during the Carboniferous and Permian periods in marine habitats. The family had a cosmopolitan distribution.
Echinoconchus is an extinct genus of brachiopods which lived during the Lower Carboniferous period. The genus was abundant and had a cosmopolitan distribution.
Echinaria is an extinct genus of brachiopods which lived during the Upper Carboniferous and Lower Permian periods. Its fossils have been found in Eurasia, North America and northern South America.
Juresaniinae is an extinct subfamily of brachiopods which lived during the Carboniferous and Permian periods in marine habitats.
Pustulinae is an extinct subfamily of medium to large-sized brachiopods which lived during the Early Carboniferous period in marine habitats. It is named after the type genus Pustula.
Antiquatonia is an extinct genus of brachiopod belonging to the order Productida and family Productidae. Specimens have been found in Carboniferous beds across many continents, suggesting the genus had a cosmopolitan distribution. Species level taxonomy of Antiquatonia is in need of revision.
Pustula is an extinct genus of brachiopods which lived during the Carboniferous period. It is the type genus of the subfamily Pustulinae. Its fossils have been found in Eurasia, North America and northern Africa.
Pulchratia is an extinct genus of brachiopods which lived in marine habitats during the Upper Carboniferous period. Its fossils have been found in North America.