Psychoda alternata | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Diptera |
Family: | Psychodidae |
Genus: | Psychoda |
Species: | P. alternata |
Binomial name | |
Psychoda alternata Say, 1824 | |
Synonyms [1] | |
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Psychoda alternata is a species of moth fly in the family Psychodidae, [1] [2] [3] [4] commonly known as the trickling filter fly [5] [6] or drain fly. [7] The larva is semiaquatic and lives in the gelatinous ooze associated with leaks of sewage effluent, [5] drains, and in trickling filter systems.
The adult female fly is about 4 mm (0.2 in) long, the male being slightly smaller. The wings are broad and held at an angle away from the body. They are clad with dense tufts of hairlike scales along the veins, and are grey, with a mottled appearance. The wing muscles are relatively weak and the insects mostly run or hop, occasionally making short flights. The males hatch first and only survive for a few days, while unmated females may live for a week. [7]
Originating in North America, this insect has spread around the world. Its range includes Europe, western Asia and South America. It had become established in Britain by 2000, in Brazil by 2006, in Norway by 2011, in Iraq and Croatia by 2013 and in Spain by 2016. [7] The larvae develop in moist areas rich in organic matter such as drains and trickling filter systems, but also in ditches and sludges of decaying organic matter. The adult insects seldom move far from where they were hatched. [6]
The eggs are laid in the moist places in which the larvae will feed. Up to 100 are laid by the female, sometimes singly, but usually in gelatinous batches of 15 to 40. The eggs are translucent and under a millimetre in diameter. They hatch in about two days at 70 °F (21 °C). The larvae are slender, whitish, cream or pale brown, with a head with strong jaws, 11 body segments, and siphons on the hindermost two segments. The larval stage lasts for about 15 days at 70 °F, and it is in the larval stage that the insect will usually overwinter, in diapause. The pupae are yellowish brown to brown and have a pair of ear-like respiratory processes on the head end. The pupal stage lasts for one or two days. Both larvae and pupae are usually to be found in the top 2.5 cm (1 in) of the substrate, sometimes deeper than this when the material is fairly dry. [6] [7]
A trickling filter system is a form of biological treatment widely used to process sewage. The sewage or other wastewater flows downwards over a medium such as gravel which is supported on a permeable membrane. In the medium live over 200 species of bacteria, algae, worms, protozoa, and insects including larvae of the drain fly; together these organisms form a biofilm which processes the fluid passing through the filter system and removes pollutants. [7]
A pupa is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages thereof being egg, larva, pupa, and imago. The processes of entering and completing the pupal stage are controlled by the insect's hormones, especially juvenile hormone, prothoracicotropic hormone, and ecdysone. The act of becoming a pupa is called pupation, and the act of emerging from the pupal case is called eclosion or emergence.
The caddisflies, or order Trichoptera, are a group of insects with aquatic larvae and terrestrial adults. There are approximately 14,500 described species, most of which can be divided into the suborders Integripalpia and Annulipalpia on the basis of the adult mouthparts. Integripalpian larvae construct a portable casing to protect themselves as they move around looking for food, while Annulipalpian larvae make themselves a fixed retreat in which they remain, waiting for food to come to them. The affinities of the small third suborder Spicipalpia are unclear, and molecular analysis suggests it may not be monophyletic. Also called sedge-flies or rail-flies, the adults are small moth-like insects with two pairs of hairy membranous wings. They are closely related to the Lepidoptera which have scales on their wings; the two orders together form the superorder Amphiesmenoptera.
Holometabolism, also called complete metamorphosis, is a form of insect development which includes four life stages: egg, larva, pupa, and imago. Holometabolism is a synapomorphic trait of all insects in the superorder Endopterygota. Immature stages of holometabolous insects are very different from the mature stage. In some species the holometabolous life cycle prevents larvae from competing with adults because they inhabit different ecological niches. The morphology and behavior of each stage are adapted for different activities. For example, larval traits maximize feeding, growth, and development, while adult traits enable dispersal, mating, and egg laying. Some species of holometabolous insects protect and feed their offspring. Other insect developmental strategies include ametabolism and hemimetabolism.
The Phoridae are a family of small, hump-backed flies resembling fruit flies. Phorid flies can often be identified by their escape habit of running rapidly across a surface rather than taking to the wing. This behaviour is a source of one of their alternate names, scuttle fly. Another vernacular name, coffin fly, refers to Conicera tibialis. About 4,000 species are known in 230 genera. The most well-known species is cosmopolitan Megaselia scalaris. At 0.4 mm in length, the world's smallest fly is the phorid Euryplatea nanaknihali.
The common green bottle fly is a blowfly found in most areas of the world and is the most well-known of the numerous green bottle fly species. Its body is 10–14 mm (0.39–0.55 in) in length – slightly larger than a house fly – and has brilliant, metallic, blue-green or golden coloration with black markings. It has short, sparse, black bristles (setae) and three cross-grooves on the thorax. The wings are clear with light brown veins, and the legs and antennae are black. The larvae of the fly may be used for maggot therapy, are commonly used in forensic entomology, and can be the cause of myiasis in livestock and pets. The common green bottle fly emerges in the spring for mating.
Psychodidae, called drain flies, sink flies, filter flies, sewer flies, or sewer gnats, is a family of true flies. Some genera have short, hairy bodies and wings giving them a "furry" moth-like appearance, hence one of their common names, moth flies. Members of the sub-family Phlebotominae which are hematophagous may be called sand flies in some countries, although this term is also used for other unrelated flies.
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A trickling filter is a type of wastewater treatment system. It consists of a fixed bed of rocks, coke, gravel, slag, polyurethane foam, sphagnum peat moss, ceramic, or plastic media over which sewage or other wastewater flows downward and causes a layer of microbial slime (biofilm) to grow, covering the bed of media. Aerobic conditions are maintained by splashing, diffusion, and either by forced-air flowing through the bed or natural convection of air if the filter medium is porous. The treatment of sewage or other wastewater with trickling filters is among the oldest and most well characterized treatment technologies.
Most organisms involved in water purification originate from the waste, wastewater or water stream itself or arrive as resting spore of some form from the atmosphere. In a very few cases, mostly associated with constructed wetlands, specific organisms are planted to maximise the efficiency of the process.
Home-stored product entomology is the study of insects which infest foodstuffs stored in the home. It deals with the prevention, detection and eradication of the pests. The five major categories of insects considered in this article are flour beetles, the drugstore beetle, the sawtoothed grain beetle, the Indianmeal moth and fruit flies.
The housefly is a fly of the suborder Cyclorrhapha. It is believed to have evolved in the Cenozoic Era, possibly in the Middle East, and has spread all over the world as a commensal of humans. It is the most common fly species found in houses. Adults are gray to black, with four dark, longitudinal lines on the thorax, slightly hairy bodies, and a single pair of membranous wings. They have red eyes, set farther apart in the slightly larger female.
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Diptera is an order of winged insects commonly known as flies. Diptera, which are one of the most successful groups of organisms on Earth, are very diverse biologically. None are truly marine but they occupy virtually every terrestrial niche. Many have co-evolved in association with plants and animals. The Diptera are a very significant group in the decomposition and degeneration of plant and animal matter, are instrumental in the breakdown and release of nutrients back into the soil, and whose larvae supplement the diet of higher agrarian organisms. They are also an important component in food chains.
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Clogmia albipunctata is a species of drain fly, a member of the family Psychodidae commonly known as the bathroom moth midge, bathroom moth fly or drain fly.
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