Reptiles, as most commonly defined, are the animals in the class Reptilia, a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates and rhynchocephalians (tuatara). As of March 2022, the Reptile Database includes about 11,700 species. In the traditional Linnaean classification system, birds are considered a separate class to reptiles. However, crocodilians are more closely related to birds than they are to other living reptiles, and so modern cladistic classification systems include birds within Reptilia, redefining the term as a clade. Other cladistic definitions abandon the term reptile altogether in favor of the clade Sauropsida, which refers to all amniotes more closely related to modern reptiles than to mammals. The study of the traditional reptile orders, customarily in combination with the study of modern amphibians, is called herpetology.
Amniotes are a clade of tetrapod vertebrates that comprises sauropsids and synapsids. They are distinguished from the other tetrapod clade — the amphibians — by the development of three extraembryonic membranes, thicker and more keratinized skin, and costal respiration.
Theropoda, whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally carnivorous, although a number of theropod groups evolved to become herbivores and omnivores. Theropods first appeared during the Carnian age of the late Triassic period 231.4 million years ago (Ma) and included all the large terrestrial carnivores from the Early Jurassic until at least the close of the Cretaceous, about 66 Ma. In the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by about 10,500 living species.
Archosauria is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term, which includes birds. Extinct archosaurs include non-avian dinosaurs, pterosaurs, and extinct relatives of crocodilians. Modern paleontologists define Archosauria as a crown group that includes the most recent common ancestor of living birds and crocodilians, and all of its descendants. The base of Archosauria splits into two clades: Pseudosuchia, which includes crocodilians and their extinct relatives, and Avemetatarsalia, which includes birds and their extinct relatives.
Archosauromorpha is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs rather than lepidosaurs. Archosauromorphs first appeared during the late Middle Permian or Late Permian, though they became much more common and diverse during the Triassic period.
Hypselosaurus is a dubious genus of titanosaurian sauropod that lived in southern France during the Late Cretaceous, approximately 70 million years ago in the early Maastrichtian. Hypselosaurus was first described in 1846, but was not formally named until 1869, when Phillip Matheron named it under the binomial Hypselosaurus priscus. The holotype specimen includes a partial hindlimb and a pair of caudal vertebrae, and two eggshell fragments were found alongside these bones. Because of the proximity of these eggshells to the fossil remains, many later authors, including Matheron and Paul Gervais, have assigned several eggs from the same region of France all to Hypselosaurus, although the variation and differences between these eggs suggest that they do not all belong to the same taxon. Hypselosaurus has been found in the same formation as the dromaeosaurids Variraptor and Pyroraptor, the ornithopod Rhabdodon, and the ankylosaurian Rhodanosaurus, as well as indeterminate bones from other groups.
Dinosaur eggs are the organic vessels in which a dinosaur embryo develops. When the first scientifically documented remains of non-avian dinosaurs were being described in England during the 1820s, it was presumed that dinosaurs had laid eggs because they were reptiles. In 1859, the first scientifically documented dinosaur egg fossils were discovered in France by Jean-Jacques Poech, although they were mistaken for giant bird eggs.
Macroolithus is an oogenus of dinosaur egg belonging to the oofamily Elongatoolithidae. The type oospecies, M. rugustus, was originally described under the now-defunct oogenus name Oolithes. Three other oospecies are known: M. yaotunensis, M. mutabilis, and M. lashuyuanensis. They are relatively large, elongated eggs with a two-layered eggshell. Their nests consist of large, concentric rings of paired eggs. There is evidence of blue-green pigmentation in its shell, which may have helped camouflage the nests.
Continuoolithus is an oogenus of dinosaur egg found in the late Cretaceous of North America. It is most commonly known from the late Campanian of Alberta and Montana, but specimens have also been found dating to the older Santonian and the younger Maastrichtian. It was laid by an unknown type of theropod. These small eggs are similar to the eggs of oviraptorid dinosaurs, but have a distinctive type of ornamentation.
Reptiles arose about 320 million years ago during the Carboniferous period. Reptiles, in the traditional sense of the term, are defined as animals that have scales or scutes, lay land-based hard-shelled eggs, and possess ectothermic metabolisms. So defined, the group is paraphyletic, excluding endothermic animals like birds that are descended from early traditionally-defined reptiles. A definition in accordance with phylogenetic nomenclature, which rejects paraphyletic groups, includes birds while excluding mammals and their synapsid ancestors. So defined, Reptilia is identical to Sauropsida.
Paleontology in Montana refers to paleontological research occurring within or conducted by people from the U.S. state of Montana. The fossil record in Montana stretches all the way back to the Precambrian. During the Late Precambrian, western Montana was covered by a warm, shallow sea where local bacteria formed stromatolites and bottom-dwelling marine life left tracks on the sediment that would later fossilize. This sea remained in place during the early Paleozoic, although withdrew during the Silurian and Early Devonian, leaving a gap in the local rock record until its return. This sea was home to creatures including brachiopods, conodonts, crinoids, fish, and trilobites. During the Carboniferous the state was home to an unusual cartilaginous fish fauna. Later in the Paleozoic the sea began to withdraw, but with a brief return during the Permian.
Paleontology in Colorado refers to paleontological research occurring within or conducted by people from the U.S. state of Colorado. The geologic column of Colorado spans about one third of Earth's history. Fossils can be found almost everywhere in the state but are not evenly distributed among all the ages of the state's rocks. During the early Paleozoic, Colorado was covered by a warm shallow sea that would come to be home to creatures like brachiopods, conodonts, ostracoderms, sharks and trilobites. This sea withdrew from the state between the Silurian and early Devonian leaving a gap in the local rock record. It returned during the Carboniferous. Areas of the state not submerged were richly vegetated and inhabited by amphibians that left behind footprints that would later fossilize. During the Permian, the sea withdrew and alluvial fans and sand dunes spread across the state. Many trace fossils are known from these deposits.
Egg fossils are the fossilized remains of eggs laid by ancient animals. As evidence of the physiological processes of an animal, egg fossils are considered a type of trace fossil. Under rare circumstances a fossil egg may preserve the remains of the once-developing embryo inside, in which case it also contains body fossils. A wide variety of different animal groups laid eggs that are now preserved in the fossil record beginning in the Paleozoic. Examples include invertebrates like ammonoids as well as vertebrates like fishes, possible amphibians, and reptiles. The latter group includes the many dinosaur eggs that have been recovered from Mesozoic strata. Since the organism responsible for laying any given egg fossil is frequently unknown, scientists classify eggs using a parallel system of taxonomy separate from but modeled after the Linnaean system. This "parataxonomy" is called veterovata.
Sankofa is an oogenus of prismatoolithid egg. They are fairly small, smooth-shelled, and asymmetrical. Sankofa may represent the fossilized eggs of a transitional species between non-avian theropods and birds.
This timeline of egg fossils research is a chronologically ordered list of important discoveries, controversies of interpretation, taxonomic revisions, and cultural portrayals of egg fossils. Humans have encountered egg fossils for thousands of years. In Stone Age Mongolia, local peoples fashioned fossil dinosaur eggshell into jewelry. In the Americas, fossil eggs may have inspired Navajo creation myths about the human theft of a primordial water monster's egg. Nevertheless, the scientific study of fossil eggs began much later. As reptiles, dinosaurs were presumed to have laid eggs from the 1820s on, when their first scientifically documented remains were being described in England. In 1859, the first scientifically documented dinosaur egg fossils were discovered in southern France by a Catholic priest and amateur naturalist named Father Jean-Jacques Poech, however he thought they were laid by giant birds.
Egg paleopathology is the study of evidence for illness, injury, and deformity in fossilized eggs. A variety of pathological conditions afflicting eggs have been documented in the fossil record. Examples include eggshell of abnormal thickness and fossil eggs with multiple layers of eggshell. The identification of egg paleopathologies is complicated by the fact that even healthy eggs can be modified during or after fossilization. Paleontologists can use techniques like cathodoluminescence or thin sectioning to identify true paleopathologies in fossil eggs. Despite the diversity of paleopathologies known from fossil eggs, the vast majority of conditions known to afflict modern eggs have not yet been seen among fossils.
Egg taphonomy is the study of the decomposition and fossilization of eggs. The processes of egg taphonomy begin when the egg either hatches or dies. Eggshell fragments are robust and can often travel great distances before burial. More complete egg specimens gradually begin to fill with sediment, which hardens as minerals precipitate out of water percolating through pores or cracks in the shell. Throughout the fossilization process the calcium carbonate composing the eggshell generally remains unchanged, allowing scientists to study its original structure. However, egg fossils buried under sediments at great depth can be subjected to heat, pressure and chemical processes that can alter the structure of its shell through a process called diagenesis.
Elongatoolithidae is an oofamily of fossil eggs, representing the eggs of oviraptorosaurs. They are known for their highly elongated shape. Elongatoolithids have been found in Europe, Asia, and both North and South America.
Dictyoolithidae is an oofamily of dinosaur eggs which have a distinctive reticulate organization of their eggshell units. They are so far known only from Cretaceous formations in China.
Plagioolithus is an oogenus of fossil egg. It is from the Early Cretaceous of Japan. It was probably laid by a bird, making it the oldest known fossil bird egg.
