Sphenodus Temporal range: | |
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Skeleton of Sphenodus nitidus (SMNS 96844-7) from the Late Jurassic of Germany | |
Tooth of Sphenodus nitidus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Chondrichthyes |
Subclass: | Elasmobranchii |
Family: | † Orthacodontidae |
Genus: | † Sphenodus Agassiz, 1843 |
Type species | |
Lamna (Sphenodus) longidens Agassiz, 1843 |
Sphenodus is an extinct genus of shark. It is placed as a member of the extinct family Orthacodontidae, which is either considered to be a member of the extinct order Synechodontiformes, [1] or the modern shark order Hexanchiformes. [2] 29 species have been described, [3] though some of these are likely synonyms, which span from the Early Jurassic (Sinemurian) to Paleocene (Danian). [2] Most species are only known from isolated teeth, [2] though the species Sphenodus macer and Sphenodus nitidus from the Late Jurassic of Germany are known from skeletons. These suggest that it was relatively large, with a body length of 2–3 metres (6.6–9.8 ft), with a fusiform body with a single dorsal fin placed posteriorly without a fin spine. [1] The teeth of Sphenodus consist of a single long, narrow central cusp, with much smaller lateral cusplets. [2] Species of Sphenodus are thought to have been actively swimming predators. [1]
The bullhead sharks are members of the genus Heterodontus, the only members of the family Heterodontidae and only living members of the order Heterodontiformes. All are relatively small, with the largest species reaching just 1.65 metres (5.5 ft) in maximum length. They are bottom feeders in tropical and subtropical waters.
The Hexanchiformes are a primitive order of sharks, numbering just seven extant species in two families. Fossil sharks that were apparently very similar to modern sevengill species are known from Jurassic specimens.
The Lamniformes are an order of sharks commonly known as mackerel sharks. It includes some of the most familiar species of sharks, such as the great white as well as less familiar ones, such as the goblin shark and megamouth shark.
Chimaeras are cartilaginous fish in the order Chimaeriformes, known informally as ghost sharks, rat fish, spookfish, or rabbit fish; the last three names are not to be confused with rattails, Opisthoproctidae, or Siganidae, respectively.
Elasmobranchii is a subclass of Chondrichthyes or cartilaginous fish, including modern sharks, rays, skates, and sawfish. Members of this subclass are characterised by having five to seven pairs of gill clefts opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil.
Rajiformes is one of the four orders in the clade Batomorphi, often referred to as the superorder Batoidea, flattened cartilaginous fishes related to sharks. Rajiforms are distinguished by the presence of greatly enlarged pectoral fins, which reach as far forward as the sides of the head, with a generally flattened body. The undulatory pectoral fin motion diagnostic to this taxon is known as rajiform locomotion. The eyes and spiracles are located on the upper surface of the head and the gill slits are on the underside of the body. Most species give birth to live young, although some lay eggs enclosed in a horny capsule.
Hybodus is an extinct genus of hybodont that lived from the Middle Triassic to the Late Cretaceous periods. Species closely related to the type species Hybodus reticulatus lived during the Early Jurassic epoch. Numerous species have been assigned to Hybodus spanning a large period of time, and it is currently considered a wastebasket taxon that is 'broadly polyphyletic' and requires reexamination.
Ptychodus is a genus of extinct large durophagous (shell-crushing) lamniform sharks from the Cretaceous period, spanning from the Albian to the Campanian. Fossils of Ptychodus teeth are found in many Late Cretaceous marine sediments worldwide.
Hybodontiformes, commonly called hybodonts, are an extinct group of shark-like cartilaginous fish (chondrichthyans) which existed from the late Devonian to the Late Cretaceous. Hybodonts share a close common ancestry with modern sharks and rays (Neoselachii) as part of the clade Euselachii. They are distinguished from other chondrichthyans by their distinctive fin spines and cephalic spines present on the heads of males. An ecologically diverse group, they were abundant in marine and freshwater environments during the late Paleozoic and early Mesozoic, but were rare in open marine environments by the end of the Jurassic, having been largely replaced by modern sharks, though they were still common in freshwater and marginal marine habitats. They survived until the end of the Cretaceous, before going extinct.
Paraorthacodus is an extinct genus of shark. It a member of the family Paraorthacodontidae, which is either placed in Hexanchiformes or in Synechodontiformes. It is known from over a dozen named species spanning from the Early Jurassic to the Paleocene, or possibly Eocene. Almost all members of the genus are exclusively known from isolated teeth, with the exception of P. jurensis from the Late Jurassic-Early Cretaceous of Europe, which is known from full body fossils from the Late Jurassic of Germany, which suggest that juveniles had a robust body with a round head, while adults had large body sizes with a fusiform profile. There was only a single dorsal fin towards the back of the body without a fin spine. The dentition had teeth with a single large central cusp along with shorter lateral cusplets, which were designed for clutching. The teeth are distinguished from those of Synechodus by the lateral cusplets decreasing in size linearly away from the central cusp rather than exponentially as in Synechodus.
Macrourogaleus is a genus of shark from the Late Jurassic Solnhofen Limestone. It is closely related to Paraorthacodus, and placed in the same family Paraorthacodontidae, The body is slender, with a single dorsal fin and a elongate caudal fin. The anal fin is also very elongate. The pectoral and pelvic fins are noticeably rounded.
Galeomorphii is a superorder of sharks. They are sometimes called galea or galean sharks. There are about 300 living species in 23 families. Galean sharks are divided into four orders: the Heterodontiformes, Orectolobiformes, Lamniformes, and Carcharhiniformes. The extinct, enigmatic Synechodontiformes are presently placed by some authors in the galeomorphs, but their taxonomic position still remains uncertain.
Orthacodontidae is an extinct family of sharks. It is disputed as to whether it belongs to the modern shark order Hexanchiformes, or the extinct order Synechodontiformes. It contains two genera. Some other authors included it in Lamniformes.
Synechodontiformes is an extinct order of prehistoric shark-like cartilaginous fish, known from the Permian to the Paleogene. They are considered to be members of Neoselachii, the group that contains modern sharks and rays.
Paracestracion is an extinct genus of heterodontiform sharks from Early Jurassic to Early Cretaceous-aged rocks of England, France, Germany and Luxembourg. The genus was first described in 1911 by Ernst Hermann Friedrich von Koken in Karl Alfred von Zittel.
Synechodus is an extinct genus of shark belonging to the family Palaeospinacidae and order Synechodontiformes. It is known from 16 species primarily spanning from the Late Triassic to Paleocene. The dentition is multicusped and was used for grasping. Several species are known from skeletal remains, including the species Synechodus ungeri from the Late Jurassic of Germany, which shows that it was relatively short with large pectoral fins and a proportionally large head with a round snout. This species is suggested to have reached a body length of 30–40 centimetres (0.98–1.31 ft). Skeletal remains are also known of the species Synechodus dubrisiensis from the Cretaceous of Europe. A skeleton of an indeterminate species is also known from the Early Cretaceous (Albian) of France, with a body length of around 70 centimetres (2.3 ft). Synechodus is suggested to have had two dorsal fins that lacked fin spines, though the number of dorsal fins is unknown in Synechodus ungeri.
Agaleus is an extinct genus of stem-galeomorph shark from the Early Jurassic Epoch. The genus Agaleus is monotypic, consisting solely of the species Agaleus dorsetensis. This species is currently only known from isolated teeth. It is the oldest known unambiguous crown group shark.
Pseudorhina is an extinct genus of squatiniform shark closely related to modern angelsharks. Fossils are known from the Late Jurassic and Early Cretaceous of Europe.
Palaeocarcharias is an extinct genus of shark, known from the Jurassic of Europe. It has only a single named species, Palaeocarcharias stromeri, which is known from exceptionally preserved specimens from the Late Jurassic (Tithonian) of Germany and France. Isolated teeth of indeterminate species from England extend the range of the genus back to the Middle Jurassic (Bathonian). Morphologically, it closely resembles carpet sharks (Orectolobiformes), and is around 1 metre (3.3 ft) in total body length. However, it shares greater similarities in tooth development with mackerel sharks (Lamniformes), including the absence of orthodentine, and has been suggested to the earliest known member of the Lamniformes or a member of a sister group to the Lamniformes. A 2018 study suggested that should be classified as the sole member of the order Palaeocarchariiformes, but a subsequent 2023 study questioned this and favoured placement in Lamniformes.
Egertonodus is an extinct genus of shark-like hybodont fish. It includes E. basanus from the Jurassic of Europe and North Africa and Cretaceous of North America, North Africa and Europe, and E. duffini from the Middle Jurassic of England. Indeterminate remains of the genus have been reported from the Early Cretaceous of Asia. E. basanus is known from preserved skull material, while E. duffini is only known from teeth. The genus is distinguished from Hybodussensu stricto by characters of the skull and teeth. E. basanus, the most common species, is thought to have reached 1.5 m in length. E. fraasi from the Late Jurassic of Germany, known from a poorly preserved full body fossil, was placed in Egertonodus in one study, but this has been subsequently questioned by other authors, due to strong differences in tooth morphology from the type species. Fossils have been found in freshwater and lagoonal environments.
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