Stelladens Temporal range: Late Cretaceous, | |
---|---|
Holotype teeth | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | Squamata |
Clade: | † Mosasauria |
Family: | † Mosasauridae |
Subfamily: | † Mosasaurinae |
Genus: | † Stelladens Longrich et al., 2023 |
Species: | †S. mysteriosus |
Binomial name | |
†Stelladens mysteriosus Longrich et al., 2023 | |
Stelladens (meaning "star tooth") is an extinct genus of mosasaurine mosasaurs from the Late Cretaceous Ouled Abdoun Basin of Morocco. The genus contains a single species, S. mysteriosus. [1]
The Stelladens holotype specimen, MHNM.KHG.1436, was discovered in the Sidi Chennane phosphate mine (Lower Couche III) of Oulad Abdoun Basin in Khouribga Province, Morocco. The specimen consists of a partial left dentary and two associated teeth, which is estimated to belong to an individual around 5 metres (16 ft) in total body length. [1]
In 2024, Longrich et al. described Stelladens mysteriosus as a new genus and species of mosasaurs based on these fossil remains. The generic name, Stelladens, combines the Latin words stella, meaning "star" and dens, meaning "tooth", in reference to the shape of the teeth. The specific name, mysteriosus, derives from the Latin mysterium, meaning "mystery", referencing the unusual tooth morphology. [1]
Undescribed teeth potentially referable to Stelladens have also been reported from the younger Upper Couche III in the Sidi Chennane locality. [2]
In 2024, Sharpe et al. argued that tooth-based holotypes generally lack sufficient character and variation data to be properly diagnostic. [3]
Mosasaurs are an extinct group of large aquatic reptiles within the family Mosasauridae that lived during the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. They belong to the order Squamata, which includes lizards and snakes.
Selmasaurus is an extinct genus of marine lizard belonging to the mosasaur family. It is classified as part of the Plioplatecarpinae subfamily alongside genera like Angolasaurus and Platecarpus. Two species are known, S. russelli and S. johnsoni; both are exclusively known from Santonian deposits in the United States.
The Mosasaurinae are a subfamily of mosasaurs, a diverse group of Late Cretaceous marine squamates. Members of the subfamily are informally and collectively known as "mosasaurines" and their fossils have been recovered from every continent except for South America.
The Halisaurinae are a subfamily of mosasaurs, a group of Late Cretaceous marine lizards. They were small to medium-sized, ranging from just under 3 meters in Eonatator sternbergi to as much as 8 or 9 meters in Pluridens serpentis. They tended to have relatively slender jaws and small, numerous teeth, suggesting a diet of small fish and other prey. Although the skeleton is primitive compared to other Mosasauridae in many respects, halisaurines had the distinctive hypocercal tail of other mosasaurids suggesting good swimming ability, and they persisted alongside other mosasaurs until the end of the Cretaceous. The earliest known remains of halisaurines occur in rocks of Santonian age and the subfamily persists until the latest Maastrichtian. Halisaurines are known from North and South America, Europe, Asia and Africa, indicating a more or less global distribution in the Late Cretaceous. Four genera are currently recognized: Eonatator, Halisaurus, Phosphorosaurus and Pluridens.
Globidens is an extinct genus of mosasaurid oceanic lizard classified as part of the Globidensini tribe in the Mosasaurinae subfamily. Globidens belongs to the family Mosasauridae, which consists of several genera of predatory marine lizards of various sizes that were prevalent during the Late Cretaceous. Specimens of Globidens have been discovered in Angola, Brazil, Morocco, Syria and the United States. Among mosasaurs, Globidens is probably most well known for the highly rounded, globe-like teeth that give it its name.
Phosphatodraco is a genus of azhdarchid pterosaur that lived during the Late Cretaceous of what is now Morocco. In 2000, a pterosaur specimen consisting of five cervical (neck) vertebrae was discovered in the Ouled Abdoun Phosphatic Basin. The specimen was made the holotype of the new genus and species Phosphatodraco mauritanicus in 2003; the genus name means "dragon from the phosphates", and the specific name refers to the region of Mauretania. Phosphatodraco was the first Late Cretaceous pterosaur known from North Africa, and the second pterosaur genus described from Morocco. It is one of the only known azhdarchids preserving a relatively complete neck, and was one of the last known pterosaurs. Additional cervical vertebrae have since been assigned to the genus, and it has been suggested that fossils of the pterosaur Tethydraco represent wing elements of Phosphatodraco.
Carinodens is an extinct genus of Cretaceous marine lizard belonging to the mosasaur family. "Carinodens" means "keel teeth" and was named in 1969 as a replacement name for Compressidens, "compressed teeth", which was already in use for a gadilidan scaphopod mollusk.
Pluridens is an extinct genus of marine lizard belonging to the Mosasauridae. Pluridens is placed in the subfamily Halisaurinae with the genera Phosphorosaurus, Eonatator and Halisaurus. Compared to related halisaurines, Pluridens had longer jaws with more teeth, and smaller eyes. It also grew large size, measuring 5–6 m (16–20 ft) long and perhaps over 9 m (30 ft) in some individuals. The jaws in some specimens are robust, and sometimes show injuries suggestive of combat. The jaws may have been used for fighting over mates or territories.
Liodon is a dubious genus of mosasaur from the Late Cretaceous, known from fragmentary fossils discovered in St James' Pit, England and possibly also the Ouled Abdoun Basin of Morocco. Though dubious and of uncertain phylogenetic affinities, Liodon was historically a highly important taxon in mosasaur systematics, being one of the genera on which the family Mosasauridae was based.
Eremiasaurus is a genus of mosasaurs, an extinct group of marine reptiles. It lived during the Maastrichtian stage of the Late Cretaceous in what is now North Africa. Only one species is known, E. heterodontus, described in 2012 from two remarkably complete fossil specimens discovered in the Ouled Abdoun Basin, Morocco. This site is known to have delivered a significant number of other related mosasaurs.
The Oulad Abdoun Basin is a phosphate sedimentary basin located in Morocco, near the city of Khouribga. It is the largest in Morocco, comprising 44% of Morocco's phosphate reserves, and at least 26.8 billion tons of phosphate. It is also known as an important site for vertebrate fossils, with deposits ranging from the Late Cretaceous (Cenomanian-Turonian) to the Eocene epoch (Ypresian), a period of about 25 million years.
Chenanisaurus is a genus of predatory abelisaurid dinosaur, with a single known species C. barbaricus. It comes from the upper Maastrichtian phosphates of the Ouled Abdoun Basin in Morocco, North Africa. The animal is known from a holotype, consisting of a partial jaw bone, and several isolated teeth found in the same beds. Chenanisaurus is one of the largest members of the Abelisauridae, and one of the last, being a contemporary of the North American Tyrannosaurus. It would have been among the dinosaur species wiped out by the Chicxulub asteroid impact and the Cretaceous-Paleogene mass extinction that followed.
Tethydraco is a genus of pterodactyloid pterosaur that lived during the Late Cretaceous period of what is now the area of present Morocco, about 66 million years ago. Tethydraco was originally assigned to the family Pteranodontidae. Some researchers argued that subsequently described material suggests that it may have been an azhdarchid, and possibly synonymous with Phosphatodraco, though this has been disputed. The type and only species is T. regalis.
Ajnabia is a genus of lambeosaurine hadrosaur from the Late Cretaceous (Maastrichtian) of Morocco. It is the first definitive hadrosaur from Africa, and is thought to be related to European hadrosaurs like Arenysaurus. The discovery of Ajnabia came as a surprise to the paleontologists who found it, because Africa was isolated by water from the rest of the world during the Cretaceous, such that hadrosaurs were assumed to have been unable to reach the continent. Ajnabia is relatively small and similar in size to its contemporary relative Minqaria, which is estimated to have reached 3.5 metres (11 ft) in total body length. Assuming that the holotype represents an adult, Ajnabia would be one of the smallest if not the smallest known hadrosaurids.
Gavialimimus is an extinct genus of plioplatecarpine mosasaur from the Maastrichtian of Morocco and possibly Angola. It was a medium-sized mosasaur measuring around 6 metres (20 ft) in total body length.
Xenodens is a potentially dubious extinct genus of marine lizard belonging to the mosasaur family. It contains a single species, X. calminechari, which is known from Late Maastrichtian phosphate deposits in the Ouled Abdoun Basin, Morocco.
Thalassotitan is an extinct genus of large mosasaurs that lived during the late Maastrichtian of the Cretaceous period in what is now Morocco, around 67 to 66 million years ago. The only known species is T. atrox, described in 2022 from fossils discovered in the Ouled Abdoun Basin, initially identified as coming from other genera such as Mosasaurus or Prognathodon. Hypothetical Thalassotitan specimens may have been found in other corners of the world, although researchers also note the possibility that they come from distinct, related taxa. It is considered to be close to the genera Prognathodon and Gnathomortis, together forming the tribe Prognathodontini. The prognathodontines are separated from other mosasaurs based on their massive jaws and robust teeth.
Minqaria is a genus of arenysaurinin lambeosaurine hadrosaur from the Late Cretaceous (Maastrichtian) Ouled Abdoun Basin of Morocco. The genus contains a single species, M. bata, known from a partial skull.
Khinjaria is an extinct genus of plioplatecarpine mosasaurid from the Late Cretaceous Ouled Abdoun Basin of Morocco. The genus contains a single species, K. acuta, known from a partial skull and vertebra. Khinjaria was likely an apex predator in its environment, as its large body size, blade-like teeth, and unusual skull morphology would have allowed it to attack large prey animals.