Varunidae | |
---|---|
Chasmagnathus convexus | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Crustacea |
Class: | Malacostraca |
Order: | Decapoda |
Infraorder: | Brachyura |
Superfamily: | Grapsoidea |
Family: | Varunidae H. Milne-Edwards, 1853 |
Subfamilies [1] | |
The Varunidae are a family of thoracotrematan crabs. The delimitation of this family, part of the taxonomically confusing Grapsoidea, is undergoing revision. For a long time, they were placed at the rank of subfamily in the Grapsidae, but they appear to be closest to Macropthalmus and the Mictyridae, which are usually placed in the Ocypodoidea. It may thus be better to merge the latter superfamily with the Grapsoidea, retaining the latter name as it is older. [2]
That notwithstanding, the revision of the Grapsoidea (in the narrow but apparently still paraphyletic sense) is also not fully completed, as many taxa remain to be restudied. Already, several former Grapsidae genera have been moved to the Varunidae, and others are tentatively placed here pending detailed study. Among the most noteworthy of these is the Chinese mitten crab, Eriocheir sinensis. [2] The genus Xenograpsus , formerly included in the Varunidae, is now placed in its own family, the Xenograpsidae. [1]
The following genera are included: [3]
Xanthidae is a family of crabs known as gorilla crabs, mud crabs, pebble crabs or rubble crabs. Xanthid crabs are often brightly coloured and are highly poisonous, containing toxins which are not destroyed by cooking and for which no antidote is known. The toxins are similar to the tetrodotoxin and saxitoxin produced by puffer fish, and may be produced by bacteria in the genus Vibrio living in symbiosis with the crabs, mostly V. alginolyticus and V. parahaemolyticus.
Eriocheir is a genus of crabs, including the Chinese mitten crab, E. sinensis. Formerly in the family Grapsidae, it is now placed in the Varunidae.
Pinnotheres is a genus of crabs, including the pea crab. Many species formerly in Pinnotheres have been placed in new genera, such as Zaops ostreus, the oyster crab and Nepinnotheres novaezelandiae, the New Zealand pea crab. The species currently recognised in the genus Pinnotheres are:
Portunidae is a family of crabs which contains the swimming crabs.
Sand bubbler crabs are crabs of the genera Scopimera and Dotilla in the family Dotillidae. They are small crabs that live on sandy beaches in the tropical Indo-Pacific. They feed by filtering sand through their mouthparts, leaving behind balls of sand that are disintegrated by the incoming high tide.
The Grapsidae are a family of crabs known variously as marsh crabs, shore crabs, or talon crabs. The family has not been confirmed to form a monophyletic group and some taxa may belong in other families. They are found along the shore among rocks, in estuaries, marshes, and in some cases pelagic among drifting seaweeds and flotsam.
Macrophthalmus is a genus of crabs which are widespread across the Indo-Pacific. It contains the following species :
Parthenopidae is a family of crabs, placed in its own superfamily, Parthenopoidea. It comprises nearly 40 genera, divided into two subfamilies, with three genera incertae sedis:
Menippidae is a family of crabs of the order Decapoda.
Ozius is a genus of crabs in the family Menippidae, containing the following species:
Pilumnoidea is a superfamily of crabs, whose members were previously included in the Xanthoidea. The three families are unified by the free articulation of all the segments of the male crab's abdomen and by the form of the gonopods. The earliest fossils assigned to this group are of Eocene age.
The Plagusiidae are a family of crabs, formerly treated as a subfamily of the family Grapsidae, but have since been considered sufficiently distinct to be a family in their own right. The family Plagusiidae includes the subfamily Plagusiinae, comprising the genera Percnon and Plagusia, which constitute a widespread group of litophilic, intertidal and subtidal crabs that are notorious for their speed and their agility.
Actumnus is a genus of crabs in the family Pilumnidae. Alongside the 28 extant species, it has a fossil record extending back into the Miocene.
Palicoidea is a superfamily of crabs, comprising the two families Crossotonotidae and Palicidae. Together, they contain 13 genera, including two genera in the Palicidae known only from fossils. The two families were previously treated as two subfamilies in a Palicidae of wider circumscription.
Epialtinae is a subfamily of crabs, containing the following genera:
Pachygrapsus is a genus of small shore crabs. Recent genetic data suggest this genus to be possibly polyphyletic.
Hemigrapsus is a genus of varunid crabs comprising thirteen species native almost exclusively in the Pacific Ocean, but two have been introduced to the North Atlantic region.
Xanthodius is a genus of crabs in the family Xanthidae, containing one exclusively fossil species and the following extant species:
Euryplacidae is a family of crabs in the superfamily Goneplacoidea which consists of 14 existing genera and 31 existing species and 8 fossil genera and 15 fossil species.
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