Vermetidae | |
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A live individual of Thylacodes squamigerus in situ | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | Gastropoda |
Subclass: | Caenogastropoda |
Order: | Littorinimorpha |
Superfamily: | Vermetoidea |
Family: | Vermetidae Rafinesque |
Genera | |
See text |
The Vermetidae, the worm snails or worm shells, are a taxonomic family of small to medium-sized sea snails, marine gastropod molluscs in the clade Littorinimorpha. [1] The shells of species in the family Vermetidae are extremely irregular, and do not resemble the average snail shell, hence the common name "worm shells" or "worm snails".
These snails usually grow cemented onto a hard surface, or cemented together in colonies.
These snails do not have typical regularly coiled gastropod shells; instead, they have very irregular elongated tubular shells which are moulded to, and cemented to, a surface of attachment such as a rock or another shell. In the adult, the apertural part of the shell is usually free, with the opening directed upward. Some species have an operculum and some do not. Damaged sections of the shell can be sealed off by calcareous septa when necessary.
Some vermetids are solitary, whereas others live in colonies, partially cemented together. The shells of species within this family vary greatly and can sometimes be extremely challenging to identify.
The empty calcareous tubes of certain marine annelid tube worms, for example the Serpulidae, can sometimes be casually misidentified as empty vermetid shells, and vice versa. The difference is that vermetid shells are shiny inside and have three shell layers, whereas the annelid worm tubes are dull inside and have only two shell layers.
The Vermetidae were recognized as the only family in the superfamily Vermetoidea in the taxonomy of Bouchet & Rocroi (2005) within the clade Littorinimorpha.
The following two subfamilies were recognized in the taxonomy of Bouchet & Rocroi (2005): [1]
Family Vermetidae Rafinesque, 1815
This classification of the Vermetoidea has been somewhat controversial. Studies, based on sperm ultrastructure, [2] and on molecular data [3] clearly place it within the clade Littorinimorpha. However, there are still a number of authors [4] [5] [6] that place it within the superfamily Cerithioidea. The genera Campanile and Thylacodes form a clade that is sister to the Cerithioidea, as shown in a study by Lydeard et al. (2002). [7]
Bandel (2006) [8] established a new subfamily Laxispirinae as one of three subfamilies he recognized within the Vermetidae:
Genera within the family Vermetidae include:
Vermetinae
† Laxispirinae
Dendropomatinae
Subfamily ?
Caenogastropoda is a taxonomic subclass of molluscs in the class Gastropoda. It is a large diverse group which are mostly sea snails and other marine gastropod mollusks, but also includes some freshwater snails and some land snails. The subclass is the most diverse and ecologically successful of the gastropods.
Littorinimorpha is a large order of snails, gastropods, consisting primarily of sea snails, but also including some freshwater snails and land snails.
Pyramidellinae is a taxonomic subfamily of minute sea snails, marine gastropod mollusks in the family Pyramidellidae, the pyrams and their allies.
Neritidae, common name the nerites, is a taxonomic family of small to medium-sized saltwater and freshwater snails which have a gill and a distinctive operculum. The family Neritidae includes marine genera such as Nerita, marine and freshwater genera such as Neritina, and freshwater and brackish water genera such as Theodoxus.
Thiaridae, common name thiarids or trumpet snails, is a family of tropical freshwater snails with an operculum, aquatic gastropod mollusks in the superfamily Cerithioidea.
Thylacodes aotearoicus is a species of sea snail, a worm snail or worm shell, a marine gastropod mollusc in the family Vermetidae, the worm snails. This species was previously known as Serpulorbis aotearoicus.
The Vermetoidea, the worm snails or worm shells, are a taxonomic superfamily of small to medium-sized sea snails, marine gastropod molluscs in the order Littorinimorpha.
Litiopidae, common name litiopids, is a family of small sea snails, marine gastropod molluscs in the clade Sorbeoconcha.
Procerithiidae is a taxonomic family of sea snails, marine gastropod molluscs in the clade Sorbeoconcha.
Dendropoma anguliferum is a species of sea snail, a marine gastropod mollusk in the family Vermetidae, the worm snails or worm shells.
Dendropoma maximum is a species of sea snail, a marine gastropod mollusk in the family Vermetidae, the worm snails or worm shells.
Dendropoma mejillonensis is a species of sea snail, a marine gastropod mollusc in the family Vermetidae, the worm snails or worm shells.
Dendropoma nebulosum is a species of sea snail, a marine gastropod mollusk in the family Vermetidae, the worm snails or worm shells.
Dendropoma petraeum is a species of sea snail, a marine gastropod mollusk in the family Vermetidae, the worm snails or worm shells.
Vermetus adansonii is a species of sea snail, a marine gastropod mollusk in the family Vermetidae, the worm snails or worm shells.
Thylacodes riisei is a species of sea snail, a marine gastropod mollusk in the family Vermetidae, the worm snails or worm shells. This species was previously known as Serpulorbis riisei.
Petaloconchus is a genus of sea snails, marine gastropod mollusks in the family Vermetidae, the worm snails or worm shells.
Vermetus is a genus of sea snails, marine gastropod mollusks in the family Vermetidae, the worm snails or worm shells.
Obtortionidae is a family of sea snails, marine gastropod molluscs in the superfamily Cerithioidea, that is within the clade Cerithimorpha or in clade Sorbeoconcha.
Thylacodes is a genus of sea snails, marine gastropod mollusks in the family Vermetidae, the worm snails or worm shells. The species in this genus were previously placed in the genus Serpulorbis.
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