A mammoth is any species of the extinct elephantid genus Mammuthus. The various species of mammoth were commonly equipped with long, curved tusks. They lived from the late Miocene epoch into the Holocene about 4,000 years ago, and various species existed in Africa, Europe, Asia, and North America. Mammoths are more closely related to living Asian elephants than African elephants.
Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Proboscideans include some of the largest known land mammals. The largest land mammal of all time may have been a proboscidean; the elephant Palaeoloxodon namadicus has been estimated to be up to 5.2 m (17.1 ft) at the shoulder and may have weighed up to 22 t, surpassing the paraceratheres, the otherwise largest known land mammals, though this estimate was made based on a single fragmentary femur and is speculative. The largest extant proboscidean is the African bush elephant, with a record of size of 4 m (13.1 ft) at the shoulder and 10.4 t. In addition to their enormous size, later proboscideans are distinguished by tusks and long, muscular trunks, which were less developed or absent in early proboscideans.
A mastodon is a member of the genus Mammut, which strictly defined, was endemic to North America and lived from the late Miocene to the early Holocene. Mastodons belong to the order Proboscidea, the same order as elephants and mammoths. Mammut is the type genus of the extinct family Mammutidae, which diverged from the ancestors of modern elephants at least 25 million years ago, during the Oligocene.
Mammutidae is an extinct family of proboscideans belonging to Elephantimorpha. It is best known for the mastodons, which inhabited North America from the Late Miocene until their extinction at beginning of the Holocene, around 11,000 years ago. The earliest fossils of the group are known from the Late Oligocene of Africa, around 24 million years ago, and fossils of the group have also been found across Eurasia. The name "mastodon" derives from Greek, μαστός "nipple" and ὀδούς "tooth", referring to their characteristic teeth.
Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains some of the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some only 1 metre (3.3 ft) in height, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.
Stegodon is an extinct genus of proboscidean, related to elephants. It was originally assigned to the family Elephantidae along with modern elephants but is now placed in the extinct family Stegodontidae. Like elephants, Stegodon had teeth with plate-like lophs that are different from those of more primitive proboscideans like gomphotheres and mammutids. The oldest fossils of the genus are found in Late Miocene strata in Asia, likely originating from the more archaic Stegolophodon, subsequently migrating into Africa. While the genus became extinct in Africa during the Pliocene, Stegodon remained widespread in South, Southeast and East Asia until the end of the Pleistocene.
Palaeoloxodon recki, often known by the synonym Elephas recki is an extinct species of elephant native to Africa and West Asia from the Pliocene or Early Pleistocene to the Middle Pleistocene. During most of its existence, the species represented the dominant elephant species in East Africa. The species is divided into five roughly chronologically successive subspecies. While the type and latest subspecies P. recki recki as well as the preceding P. recki ileretensis are widely accepted to be closely related to Eurasian Palaeoloxodon, the relationships of the other, chronologically earlier subspecies to P. recki recki and P. recki ileretensis are uncertain, with it being suggested they are unrelated and should be elevated to separate species.
The straight-tusked elephant is an extinct species of elephant that inhabited Europe and Western Asia during the Middle and Late Pleistocene. It was larger than any living elephant, with adult males suggested to reach 3.81–4.2 metres (12.5–13.8 ft) in shoulder height, and 11.3–15 tonnes in weight. Like modern elephants, the straight-tusked elephant lived in herds, flourishing during interglacial periods, when its range would extend as far north as Great Britain. Skeletons found in association with stone tools and wooden spears suggest they were scavenged and hunted by early humans, including Neanderthals. It is the ancestral species of most dwarf elephants that inhabited islands in the Mediterranean.
Gomphotherium is an extinct genus of gomphothere proboscidean from the Neogene of Eurasia, Africa and North America. The genus is probably paraphyletic.
Anancus is an extinct genus of "tetralophodont gomphothere" native to Afro-Eurasia, that lived from the Tortonian stage of the late Miocene until its extinction during the Early Pleistocene, roughly from 8.5–2 million years ago.
Mammuthus trogontherii, sometimes called the steppe mammoth, is an extinct species of mammoth that ranged over most of northern Eurasia during the Early and Middle Pleistocene, approximately 1.7 million-200,000 years ago. One of the largest mammoth species, it evolved in East Asia during the Early Pleistocene, around 1.8 million years ago, before migrating into North America around 1.5 million years ago, and into Europe during the Early/Middle Pleistocene transition, around 1 to 0.7 million years ago. It was the ancestor of the woolly mammoth and Columbian mammoth of the later Pleistocene.
Sinomastodon is an extinct gomphothere genus known from the Late Miocene to Early Pleistocene of Asia, including China, Japan, Thailand, Myanmar, Indonesia and probably Kashmir.
Palaeoloxodon namadicus is an extinct species of prehistoric elephant known from the early Middle to Late Pleistocene of the Indian subcontinent, and possibly also elsewhere in Asia. The species grew larger than any living elephant, and some authors have suggested it to have been the largest known land mammal based on extrapolation from fragmentary remains, though these estimates are speculative.
Palaeomastodon is an extinct genus within the elephant order Proboscidea. Its fossils have been extracted from Oligocene strata conventionally dated to 33.9-23.03 million years old. Usually considered an ancestor or near-ancestor of elephants or mastodons as a member of Elephantiformes it lived in marshes and fluvial-deltaic environments of what is now Egypt, Ethiopia, Libya, and Saudi Arabia.
Zygolophodon is an extinct genus of mammutid proboscidean that lived during the Miocene in Africa, Eurasia, and North America.
Choerolophodon is an extinct genus of proboscidean that lived during the Miocene of Eurasia and Africa. Fossils of Choerolophodon have been found in Africa, Southeast Europe, Turkey, Iraq, Iran, the Indian subcontinent, and China.
Elephantiformes is a suborder within the order Proboscidea. Members of this group are primitively characterised by the possession of upper tusks, an elongated mandibular symphysis and lower tusks, and the retraction of the facial region of the skull indicative of the development of a trunk. The earliest known member of the group, Dagbatitherium is known from the Eocene (Lutetian) of Togo, which is only known from isolated teeth, while other primitive elephantiforms like Phiomia and Palaeomastodon are known from the Early Oligocene onwards. Phiomia and Palaeomastodon are often collectively referred to as "palaeomastodonts" and assigned to the family Palaeomastodontidae. Most diversity of the group is placed in the subclade Elephantimorpha, which includes mastodons, as well as modern elephants and gomphotheres (Elephantida). It is disputed as to whether Phiomia is closely related to both Mammutidae and Elephantida with Palaeomastodon being more basal, or if Palaeomastodon is closely related to Mammutidae and Phiomia more closely related to Elephantida.
Amebelodontidae is an extinct family of large herbivorous proboscidean mammals related to elephants. They were formerly assigned to Gomphotheriidae, but recent authors consider them a distinct family. They are distinguished from other proboscideans by having flattened lower tusks and very elongate mandibular symphysis. The lower tusks could grow considerable size, with those of Konobelodon reaching 1.61 metres (5.3 ft) in length. Their molar teeth are typically trilophodont, and possessed posttrite conules. In the past, amebelodonts' shovel-like mandibular tusks led to them being portrayed scooping up water plants, however, dental microwear suggests that they were browsers and mixed feeders. The lower tusks have been proposed to have had a variety of functions depending on the species, including stripping bark, cutting through vegetation, as well as possibly digging. They first appeared in Africa during the Early Miocene, and subsequently dispersed into Eurasia and then North America. They became extinct by the beginning of the Pliocene. While some phylogenetic studies have recovered Amebelodontidae as a monophyletic group that forms the sister group to Gomphotheriidae proper, some authors have argued that Amebelodontidae may be polyphyletic, with it being suggested that the shovel-tusked condition arose several times independently within Gomphotheriidae, thus rendering the family invalid.
Eozygodon is an extinct genus of proboscidean in the family Mammutidae that is based on a single species, E. morotoensis, named in 1983. It is known from the Early Miocene of Africa and well as possibly the Middle Miocene of China. It is considered a primitive member of the family, retaining a long lower jaw (longirostrine) with lower tusks. The upper tusks are small and are parallel to each other. The body mass is considerably smaller than the American mastodon. Dental mesowear from East African specimens suggests a browsing diet.
The research history of Mammut is extensive given its complicated taxonomic and non-taxonomic histories, with the earliest recorded fossil finds dating back to 1705 in Claverack, New York during the colonial era of what is now the United States of America. Initially thought to belong to biblical antediluvian giants, the fossils were later determined to belong to a proboscidean species as a result of more complete 18th century finds from the locality of Big Bone Lick in what is now Kentucky. The molars were studied by European and American naturalists, who were generally baffled on its lack of analogue to modern elephants, leading to varying hypothesis on the affinities of the teeth. More complete skeletons were found after the independence of the United States colonies from Great Britain within the early 19th century. American historians of the 21st century have made arguments that the early history of M. americanum finds and studies played major roles in shaping American nationalism on the basis of the large sizes and relative completeness of the fossils to disprove the negative theory of social degeneracy in North America.
Tusks of "M." borsoni from Milia, Greece, which are the longest tusks ever found
Molar tooth
Lower jaw
