Anagyrus

For the deme of ancient Attica, see Anagyrous.

Anagyrus
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hymenoptera
Family:	Encyrtidae
Subfamily:	Tetracneminae
Genus:	Anagyrus Howard, 1896
Species
See text
Synonyms
Aagyrus [1] [2]

A vial containing A. lopezi , at a release ceremony organised and hosted by the Thai Department of Agriculture in the country's northeastern Khon Kaen province. The wasp is being used to help control cassava mealybug outbreaks in the country.

Anagyrus is a large genus of parasitic wasps from the family Encyrtidae. ^{[3] [4] [5] [6] [7] [8]} Anagyrus is distributed throughout the world. ^{[9] [10]} A subgenus of Anagyrus is known as Nesoanagyrus (Beardsley 1969) ^[11]

Species

There are at least 247 species in this genus which consists of:

• Anagyrus abatos (Noyes & Menezes, 2000) ^{[12] [13] [14] [15] [16] [17] [18] [19] [20]}
• Anagyrus abdulrassouli (Myartseva, Sugonjaev & Trjapitzin, 1982) ^{[21] [22] [23] [24] [25] [26] [27] [28] [29]}
• Anagyrus abyssinicus Compere, 1939
• Anagyrus aceris Noyes & Hayat, 1994
• Anagyrus aciculatus (Blanchard, 1940)
• Anagyrus adamsoni Timberlake, 1941
• Anagyrus aega Noyes, 2000
• Anagyrus aegyptiacus Moursi, 1948
• Anagyrus agraensis Saraswat 1975
• Anagyrus alami Hayat 1970
• Anagyrus albatus Myartseva, 1982
• Anagyrus aligarhensis Agarwal & Alam 1959
• Anagyrus almoriensis Shafee, Alam & Agarwal, 1975
• Anagyrus amnicus Prinsloo, 1985
• Anagyrus amoenus Compere, 1939
• Anagyrus amudaryensis (Myartseva, 1982)
• Anagyrus ananatis Gahan, 1949
• Anagyrus antoninae Timberlake, 1920
• Anagyrus aper Noyes & Menezes 2000
• Anagyrus aquilonaris (Noyes & Hayat 1984)
• Anagyrus arambourgi Risbec, 1955
• Anagyrus aranzadii (Mercet 1921)
• Anagyrus archangelskayae Trjapitzin, 1972
• Anagyrus arenaria Prinsloo, 1985
• Anagyrus argyrus (Burks 1952)
• Anagyrus ashkhabadensis Myartseva, 1981
• Anagyrus aurantifrons Compere, 1926
• Anagyrus australiensis (Howard, 1898)
• Anagyrus bambeyi Risbec 1951
• Anagyrus belibus (Walker, 1837)
• Anagyrus bellator (De Santis, 1972)
• Anagyrus bellus (Girault 1921)
• Anagyrus beneficians Compere, 1943
• Anagyrus bermudensis (Kerrich, 1982)
• Anagyrus bicolor Noyes & Hayat, 1994
• Anagyrus bohemanni (Westwood, 1837)
• Anagyrus bouceki Hoffer, 1953
• Anagyrus brachyclavae (Wu & Xu, 2001)
• Anagyrus brevicornis (Shamim & Shafee, 1985)
• Anagyrus brevistigma De Santis, 1964
• Anagyrus bugandaensis Compere, 1939
• Anagyrus californicus (Compere 1947)
• Anagyrus calyxtoi Noyes, 2000
• Anagyrus cepio Noyes, 2000
• Anagyrus cercides Noyes & Menezes 2000
• Anagyrus chilensis (Brèthes, 1916) ^{[30] [31]}
• Anagyrus chrysos (Noyes & Hayat, 1994) ^{[32] [33]}
  • jr synonym Aagyrus chrysos(Noyes & Hayat, 1994) ^{[34] [35] [36] [37] [38]}
• Anagyrus cilla Noyes & Menezes 2000
• Anagyrus clauseni Timberlake, 1924
• Anagyrus clavatus Sushil & Khan, 1996
• Anagyrus coccidivorus Dozier, 1932
• Anagyrus coccurae Sugonjaev, 1962
• Anagyrus comptoni Noyes & Hayat, 1994
• Anagyrus cooki (Girault 1919)
• Anagyrus coxalis Noyes & Hayat, 1994
• Anagyrus cristinae Noyes & Menezes 2000
• Anagyrus dactylopii (Howard 1898)
• Anagyrus darevskii (Trjapitzin 1965)
• Anagyrus diffinis Noyes & Hayat, 1994
• Anagyrus discolor Noyes & Hayat, 1994
• Anagyrus diversicornis (Howard 1894)
• Anagyrus dozieri Tachikawa, 1956
• Anagyrus echion Noyes & Hayat, 1994
• Anagyrus elgeri (Kerrich, 1982)
• Anagyrus elizabethae Noyes & Hayat, 1994
• Anagyrus elpis Noyes, 2000
• Anagyrus emeljanovi Trjapitzin, 1972
• Anagyrus ephyra Noyes & Hayat, 1994
• Anagyrus eudora Noyes, 2000
• Anagyrus euroto Noyes, 2000
• Anagyrus fasciiscapus (Girault 1932)
• Anagyrus fatimae Fatima & Agarwal, 1993
• Anagyrus ferus Noyes & Hayat, 1984
• Anagyrus flaviceps Timberlake, 1941
• Anagyrus flavimesopleurum (Girault 1917)
• Anagyrus floris Noyes & Hayat, 1994
• Anagyrus foersteri (Girault 1915)
• Anagyrus fraternus Perkins 1910
• Anagyrus frontolatus Sushil & Khan, 1996
• Anagyrus fujianensis Gu, 2004
• Anagyrus fujikona Tachikawa, 1963
• Anagyrus fusciventris (Girault, 1915)
• Anagyrus fuscus Shi, Si & Wang, 1994
• Anagyrus galinae (Myartseva, 1982)
• Anagyrus gaudens (Kerrich 1982)
• Anagyrus gemma Noyes, 2000
• Anagyrus gracilis (Hayat, 1970)
• Anagyrus gravis Noyes & Hayat, 1994
• Anagyrus greeni Howard 1896
• Anagyrus grenfelli Noyes & Hayat, 1994
• Anagyrus hainanensis Noyes & Hayat, 1994
• Anagyrus haloxyli Sugonjaev, 1968
• Anagyrus hammadae Trjapitzin & Rozanov, 1972
• Anagyrus hansoni Noyes & Menezes 2000
• Anagyrus haroldi Noyes & Hayat, 1994
• Anagyrus hayati Sushil & Khan, 1996
• Anagyrus hebes Noyes, 2000
• Anagyrus henanensis Shi, Si & Wang, 1994
• Anagyrus hippocoon Trjapitzin, 1965
• Anagyrus impar Noyes & Hayat, 1994
• Anagyrus indicus (Subba Rao, 1967)
• Anagyrus inermis Noyes & Hayat, 1994
• Anagyrus insolitus (Howard 1897)
• Anagyrus ishaqi Fatima, 1999
• Anagyrus jacksoni Noyes & Hayat, 1994
• Anagyrus jenniferae Noyes & Hayat, 1994
• Anagyrus juani Noyes, 2000
• Anagyrus jucundus De Santis, 1964
• Anagyrus juno Noyes & Hayat, 1994
• Anagyrus kamali Moursi 1948
• Anagyrus kivuensis Compere, 1939
• Anagyrus laeviceps Perkins, 1910
• Anagyrus lampe Noyes & Menezes 2000
• Anagyrus lebadeia Noyes, 2000
• Anagyrus levis Noyes & Hayat, 1994
• Anagyrus lineatipes (Girault 1919)
• Anagyrus livens Noyes, 2000
• Anagyrus lizanorum Noyes & Menezes 2000
• Anagyrus loecki Noyes & Menezes 2000
• Anagyrus longicornis Mercet 1923
• Anagyrus longipennis Shafee, Alam & Agarwal, 1975
• Anagyrus lopezi (De Santis, 1964)
• Anagyrus luci Noyes & Hayat, 1994
• Anagyrus lutescens Noyes & Hayat, 1994
• Anagyrus major Perkins, 1910
• Anagyrus malayensis Noyes & Hayat, 1994
• Anagyrus malenotus (De Santis, 1972)
• Anagyrus mandibularis Sushil & Khan, 1996
• Anagyrus mangicola Noyes, 1990
• Anagyrus maritzae Noyes, 2000
• Anagyrus marquesanus Timberlake, 1941
• Anagyrus matho Noyes, 2000
• Anagyrus matritensis (Mercet, 1921)
• Anagyrus mazaces Noyes & Hayat, 1994
• Anagyrus minium (Mercet, 1921)
• Anagyrus mirtesae Noyes & Menezes, 2000
• Anagyrus mirus (Girault, 1915)
• Anagyrus mirzai Agarwal & Alam 1959
• Anagyrus mohani Sushil & Khan, 1996
• Anagyrus montivagus (De Santis, 1964)
• Anagyrus mumfordi Timberlake, 1941
• Anagyrus mycale Noyes, 2000
• Anagyrus narcicius Salazar, 1981
• Anagyrus nesticoccus Dang & Wang, 2002
• Anagyrus niger (Ishii, 1928)
• Anagyrus nigrescens Compere, 1939
• Anagyrus nigricans Perkins, 1910
• Anagyrus nigriceps (De Santis, 1972)
• Anagyrus nigricorpus Shafee, Alam & Agarwal, 1975
• Anagyrus nigriflagellum (Girault 1915)
• Anagyrus nigritus (Howard, 1898)
• Anagyrus nishidai Noyes & Hayat, 1994
• Anagyrus nitidus Trjapitzin & Rzaeva, 1967
• Anagyrus novickyi Hoffer, 1953
• Anagyrus obodas Noyes & Hayat, 1994
• Anagyrus ocellatus Noyes & Menezes 2000
• Anagyrus odacon Noyes & Menezes 2000
• Anagyrus orbitalis Timberlake, 1941
• Anagyrus paralia Noyes & Menezes, 2000
• Anagyrus pergandei Dang & Wang, 2002
• Anagyrus petronae Noyes, 2000
• Anagyrus phaena Noyes & Menezes 2000
• Anagyrus phasis Noyes, 2000
• Anagyrus phya Noyes & Menezes 2000
• Anagyrus pilosus Ishii, 1928
• Anagyrus procles Noyes, 2000
• Anagyrus pseudococci (Girault 1915)
• Anagyrus pulcher (Ashmead, 1888)
• Anagyrus pulchricornis (Howard 1894)
• Anagyrus pullus Compere, 1939
• Anagyrus punctifrons Timberlake 1941
• Anagyrus putonophilus Compere 1947
• Anagyrus qadrii (Hayat, Alam & Agarwal, 1975)
• Anagyrus quadrimaculatus Xu & He 1996
• Anagyrus ranchiensis Shamim & Shafee, 1984
• Anagyrus rapo Noyes & Menezes, 2000
• Anagyrus remotor Noyes, 2000
• Anagyrus rhopaloides Noyes, 2000
• Anagyrus rifens Noyes & Menezes 2000
• Anagyrus rosichoni Noyes & Menezes 2000
• Anagyrus rotundiceps (Girault, 1932)
• Anagyrus rubellus (Annecke, 1974)
• Anagyrus rubinae Noyes & Hayat, 1994
• Anagyrus rugas Noyes & Hayat, 1994
• Anagyrus rusticus (De Santis, 1964)
• Anagyrus sabas Noyes & Hayat, 1994
• Anagyrus saccharicola Timberlake, 1932
• Anagyrus saintpierrei Girault, 1913
• Anagyrus saipanensis Doutt, 1952
• Anagyrus salazari Noyes & Menezes 2000
• Anagyrus sameenae Noyes & Hayat, 1994
• Anagyrus sawadai Ishii, 1928
• Anagyrus scaea Noyes, 2000
• Anagyrus scapularis Myartseva, 1982
• Anagyrus schoenherri (Westwood, 1837)
• Anagyrus scimitar Noyes & Hayat, 1994
• Anagyrus securicornis Domenichini, 1953
• Anagyrus semifulvus Girault, 1915
• Anagyrus shahidi Hayat, 1979
• Anagyrus siccus (Prinsloo & Annecke, 1976)
• Anagyrus similis (Girault 1915)
• Anagyrus sinensis Noyes & Hayat, 1994
• Anagyrus sinope Noyes & Menezes 2000
• Anagyrus smithi Doutt, 1952
• Anagyrus sogdianus Sugonjaev, 1968
• Anagyrus sophax Noyes & Menezes 2000
• Anagyrus spaici (Hoffer, 1970) ^{[39] [40]}
• Anagyrus spica (Girault 1921)
• Anagyrus subalbipes Ishii, 1928
• Anagyrus subflaviceps (Girault 1915)
• Anagyrus subnigricornis Ishii, 1928
• Anagyrus subproximus (Silvestri, 1915)
• Anagyrus subtilis Noyes & Hayat, 1994
• Anagyrus sucro Noyes, 2000
• Anagyrus suia Noyes, 2000
• Anagyrus surekhae Noyes & Menezes 2000
• Anagyrus swezeyi Timberlake, 1919
• Anagyrus tamaricicola Trjapitzin, 1968
• Anagyrus tanystis De Santis, 1964
• Anagyrus telon Noyes & Menezes 2000
• Anagyrus tenuis Noyes & Hayat, 1994
• Anagyrus terebratus (Howard 1894)
• Anagyrus thailandicus (Myartseva, 1979)
• Anagyrus theana Noyes, 2000
• Anagyrus theon Noyes & Hayat, 1994
• Anagyrus thoe Noyes, 2000
• Anagyrus thyridopterygis (Ashmead, 1886)
• Anagyrus tibimaculatus Agarwal, 1965
• Anagyrus townsendi (Howard, 1898)
• Anagyrus tricolor (Girault 1913)
• Anagyrus trinidadensis (Kerrich 1953)
• Anagyrus tristis Noyes & Hayat, 1994
• Anagyrus trjapitzini Sharipov, 1983
• Anagyrus tyana Noyes, 2000
• Anagyrus tymber Noyes, 2000
• Anagyrus umairi Noyes & Hayat, 1994
• Anagyrus ussuriensis Sharkov, 1984
• Anagyrus varithorax (Girault 1923)
• Anagyrus villalobosi Noyes & Menezes 2000
• Anagyrus vladimiri - mealybug biocontrol ^[41]
• Anagyrus vochos Noyes & Hayat, 1994
• Anagyrus vulso Noyes & Menezes 2000
• Anagyrus wayfoongi Noyes & Hayat, 1994
• Anagyrus xanthogaster Perkins, 1910
• Anagyrus yuccae (Coquillet, 1890)
• Anagyrus zaitzevi Trjapitzin, 1972
• Anagyrus zama Noyes & Menezes 2000
• Anagyrus zubairi Noyes & Hayat, 1994
• Anagyrus zygia (Trjapitzin, 1966)

References

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Further reading

• "Taxonomy browser (Anagyrus)". National Center for Biotechnology Information .
• Annals - Entomological Society of America. [College Park, Md., etc.]Entomological Society of America.
• Annals of the Entomological Society of America. [College Park, Md., etc.]: Entomological Society of America.
• Archiv für Naturgeschichte. Berlin: Nicolai, 1912-
• Archiv für Naturgeschichte. Berlin: Nicolai, 1835-
• Australian Hymenoptera Chalcidoidea / by A.A. Girault. Brisbane: Queensland Museum, 1912-1916.
• Bibliographia zoologica. Leipzig: Wilhelm Engelmann, 1896-1934.
• Bulletin - United States National Museum. Washington: Smithsonian Institution Press, [etc.];1877-1971.
• Bulletin / Illinois Natural History Survey. Urbana, State of Illinois, Dept. of Registration and Education, Natural History Survey Division, 1918-1985.
• Bulletin of the British Museum (Natural History). London: BM (NH) Bulletin of zoological nomenclature. London.
• Catalog of hymenoptera in America north of Mexico / prepared cooperatively by specialists on the various groups of Hymenoptera under the direction of Karl V. Krombein ... [et al.]. Washington: Smithsonian Institution Press, 1979-
• Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus / auctore, C.G. de Dalla Torre. Lipsiae: G. Engelmann, 1894.
• Classification of the chalcid flies, or the superfamily Chalcidoidea, with descriptions of new species in the Carnegie museum, collected in South America by Herbert H. Smith. By William Harris Ashmead... Pittsburg: Carnegie institute, 1904.
• Destructive insects affecting Ohio shade and forest trees / J. S. Houser. [Wooster, Ohio: Experiment Station, 1918]
• Dictionnaire universel d'histoire naturelle: résumant et complétant tous les faits présentés par les encyclopédies, les anciens dictionnaires scientifiques, les oeuvres complètes de Buffon, et les... traités spéciaux sur les diverses branches de
• Paris: Chez les editeurs MM. Renard, Martinet et cie, rue et Hotel Mignon, 2 (quartier de l'École-de-Médecine); et chez Langlois et Leclercq, rue de la Harpe, 81; Victor Masson, Place de l'Ecole-de-Médecin 1847-1849
• Dictionnaire universel d'histoire naturelle. Par Ch. d'Orbigny avec la collaboration de mm. Arago... [et al.]. Paris, A. Pilon et cie [1867?]
• Die Natugeschichte des Cajus Plinius Secundus: ins Deutsche übersetzt und mit Anmerkungen versehen / von G.C. Wittstein. Leipzig: Gressner & Schramm, 1881-1882.
• Entomological News. [Philadelphia] American Entomological Society, 1925-
• Experiment station record. Washington: G.P.O., 1889-1946.
• Fauna hawaiiensis; being the land-fauna of the Hawaiian islands. by various authors, 1899-1913. Cambridge [Eng.]: The University press, 1913.
• Fauna ibérica. por Ricardo García Mercet. Madrid: Museo Nacional de Ciencias Naturales, 1921.
• Flora oder Botanische Zeitung: welche Recensionen, Abhandlungen, Aufsätze, Neuigkeiten und Nachrichten, die Botanik betreffend, enthält /herausgegeben von der Königl. Botanischen Gesellschaft in Regensburg. Regensburg: Die Gesellschaft, 1818-1965.
• Hymenoptera of America north of Mexico: synoptic catalog, second supplement / prepared by the staff and collaborators of the Hymenoptera Unit, Insect Identification and Parasite Introduction Research Branch, Entomology Research Division, Agricultural Research Service, under the Washington, D.C.: U.S. Dept. of Agriculture, 1967.
• Index to the literature of American economic entomology. Melrose Highlands, Mass. [etc.], American Association of Economic Entomologists [etc.], 1917-62.
• Index zoologicus. An alphabetical list of names of genera and subgenera proposed for use in zoology as recorded in the "Zoological record" 1880-1900, together with other names not included in the "Nomenclator zoologicu Comp. (for the Zoological society of London) by Charles Owen Waterhouse, and ed. London, Printed for the Society, 1902.
• International Catalogue of Scientific Literature, 1901-1914. London. Published for the International Council by the Royal Society of London 1901-1920
• Johnson's Gardeners' dictionary and cultural instructor. London, A. T. De La Mare printing and publishing co., ltd.[1916]
• Jornal de sciencias mathematicas, physicas e naturaes / Academia Real das Sciencias de Lisboa. Lisboa: Academia Real das Sciencias de Lisboa, 1866-1927
• Memoirs of the Queensland Museum. Brisbane, Queensland Museum, 1912-
• Occasional papers of the California Academy of Sciences. San Francisco: California Academy of Sciences,
• Pamphlets on forestry in Ohio. [1900?-]
• Proceedings of the Entomological Society of Washington. [Washington, etc.: Entomological Society of Washington]
• Proceedings of the Hawaiian Entomological Society. [Honolulu: Hawaiian Entomological Society]
• Proceedings of the United States National Museum. Washington: Smithsonian Institution Press [etc.]
• Psyche. [Cambridge, Mass.: Cambridge Entomological Club]
• Revision of the parasitic hymenopterous insects of the genus Aphycus Mayr, with some notice of related genera. By P. H. Timberlake. Washington, Govt. Print. Off., 1916.
• The Canadian entomologist. Ottawa [etc.] Runge Press [etc.]
• The Entomologist. London: Simpkin, Marshall & Co., [1877-]
• The Gardeners' chronicle: a weekly illustrated journal of horticulture and allied subjects. London: [Gardeners Chronicle], 1874-1955.
• The Review of applied entomology. [Farnham Royal, Eng., etc.: Commonwealth Agricultural Bureaux, etc.]
• The standard cyclopedia of horticulture; a discussion, for the amateur, and the professional and commercial grower, of the kinds, characteristics and methods of cultivation of the species of plants grown in the regions of the United States and Illustrated with colored plates, four thousand engravings New York, Macmillan, 1919 [c1914]
• Transactions of the Entomological Society of London. London, The Society.
• Zoologischer Anzeiger. Jena, VEB Gustav Fischer Verlag.
• Kairomonal response of the parasitoid Anagyrus spec. nov. near pseudococci to the sex pheromone of the vine mealybug.
• Foraging behavior of the mealybug parasitoid Anagyrus sp. nov. nr. sinope (Hymenoptera: Encyrtidae).
• Progeny fitness of the mealybug parasitoid Anagyrus sp. nov. nr. Sinope (Hymenoptera: Encyrtidae) as affected by brood size, sex ratio, and host quality.
• Chinese fauna of parasitic wasps on scale insects.
• Interference of ants (Hymenoptera: Formicidae) with biological control of the vine mealybug Planococcus ficus (Signoret) (Hemiptera: Pseudococcidae).