Anakamacops Temporal range: Guadalupian, | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Amphibia |
Order: | † Temnospondyli |
Family: | † Dissorophidae |
Genus: | † Anakamacops Li and Cheng, 1999 |
Species | |
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Anakamacops (meaning "similar to Kamacops " in Greek) is a genus of dissorophid temnospondyl from the early Middle Permian of China. It is known from the right side of a snout that was described in 1999 from the Dashankou locality of the Xidagou Formation, which is within the city of Yumen. The type species was named A. petrolicus because Yumen is an oil-producing city (petrol). [1] More substantial material, including a partial skull and partial mandibles, was described by Liu (2018). [2]
Anakamacops is most similar to Kamacops, sharing features such as a choana widely separated from the interpterygoid vacuities and extensive exostosis ornamenting the skull roof. Both are large taxa; the most complete specimen of Anakamacops measures 26 cm in length despite lacking most of the snout. A number of potentially unique features were suggested based on the additional material described by Liu (2018), such as a relatively edentulous (lacking teeth) vomer and paired (rather than a single, unpaired) occipital ridge.
Dissorophidae |
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Cacops, is a genus of dissorophid temnospondyls from the Kungurian stage of the early Permian of the United States. Cacops is one of the few olsoniforms whose ontogeny is known. Cacops fossils were almost exclusively known from the Cacops Bone Bed of the Lower Permian Arroyo Formation of Texas for much of the 20th century. New material collected from the Dolese Brothers Quarry, near Richards Spur, Oklahoma in the past few decades has been recovered, painting a clearer picture of what the animal looked and acted like.
Dissorophidae is an extinct family of medium-sized, temnospondyl amphibians that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America.
Zygosaurus is an extinct genus of dissorophid temnospondyl from the Middle-Late Permian of Russia. It was described in 1848 by Eduard Eichwald, making it the first dissorophid to be described and is known from a single species, Zygosaurus lucius. The location of the holotype, and only known specimen, is unknown, and although casts are reposited in several institutions, little is known about this taxon beyond qualitative aspects of the skull. The skull was estimated to be around 20 cm in length, making it one of the largest dissorophids, being only slightly smaller than Kamacops.
Saharastega is an extinct genus of basal temnospondyl which lived during the Late Permian period, around 251 to 260 million years ago. Remains of Saharastega, discovered by paleontologist Christian A. Sidor at the Moradi Formation in Niger, were described briefly in 2005 and more comprehensively in 2006. The description is based on a skull lacking the lower jaws.
Kamacops is a genus of dissorophid temnospondyls known from the Middle to Late Permian of Russia that was described by Yuri Gubin in 1980. It is known from a single species, Kamacops acervalis, material of which is currently reposited in the Paleontological Institute of the Russian Academy of Sciences. Kamacops is one of the youngest dissorophids, along with Iratusaurus and Zygosaurus from Russia and Anakamacops from China and was one of the largest known dissorophids, with an estimated skull length of 24–30 cm. It is typically recovered as being most closely related to Zygosaurus and to the North American Cacops. A detailed study of the braincase region was performed by Schoch (1999), one of the first to examine this region in dissorophids. This study revealed extensive co-ossification of the braincase.
Konzhukovia is an amphibian genus that belongs to an extinct group of temnospondyls, the largest clade of basal tetrapods including about 198 genera, 292 species, and more than half of which were alive during the early Mesozoic period. The animal was a predator that lived about 260 million years ago, and could get up to about three meters in length. Specifically, Konzukovia lived during the Permian, between 252 and 270 million years ago according to the type of rock the fossil was found in. There are three species within this genus, K. vetusta, K. tarda, and K. sangabrielensis, the first two originating from Russia while the latest originating from Southern Brazil. The discovery of this specimen in Southern Brazil provided more evidence to support the idea that during this animals existence, there was a “biological corridor” because of the supercontinent Pangea, allowing these species to be found so far apart from each other. Konzhukovia belongs to the family Archegosauridae, a family consisted of large temnospondyls that most likely compare to modern day crocodiles. Since the discovery of the latest species, K. sangabrielensis, Pacheco proposes that there must be the creation of a new family, Konzhokoviidae, a monophyletic group in a sister-group relationship with Stereospondlyi in order to accommodate the three species. Konzhukovia skulls usually exhibit typical rhinesuchid features including an overall parabolic shape, small orbits located more posteriorly, and the pterygoids do not reach the vomer. These animals were long-snouted amphibians that had clear adaptations made for fish catching, as well as exemplifying aquatic features.
Trematopidae is a family of dissorophoid temnospondyl spanning the late Carboniferous to the early Permian. Together with Dissorophidae, the family forms Olsoniformes, a clade comprising the medium-large terrestrial dissorophoids. Trematopids are known from numerous localities in North America, primarily in New Mexico, Oklahoma, and Texas, and from the Bromacker quarry in Germany.
Anconastes is an extinct genus of dissorophoid temnospondyl within the family Trematopidae. It is known from two specimens from the Late Carboniferous Cutler Formation of north-central New Mexico in the southwestern United States. The genus name derives from two Greek roots, ankos and nastes ("inhabitant"), which refers to the type locality of El Cobre Canyon where the specimens were found. The specific name is derived from the Latin word vesperus ("western"). The more complete specimen, the holotype, is a partial skull with articulated mandibles and a substantial amount of the postcranial skeleton. The less complete specimen, the paratype, consists only of the right margin of the skull with an articulated mandible.
Ecolsonia is an extinct genus of trematopid temnospondyl. Its phylogenetic position within Olsoniformes has been historically debated, but it is presently considered to be a trematopid.
Dissorophus (DI-soh-ROH-fus) is an extinct genus of temnospondyl amphibian that lived during the Early Permian Period about 273 million years ago. Its fossils have been found in Texas and in Oklahoma in North America. Its heavy armor and robust build indicate Dissorophus was active on land, similar to other members of the clade Dissorophidae that are known from the Late Carboniferous to the Early Permian periods. Dissorphus is distinguished by its small body size, disproportionately large head and short trunk.
Tersomius is an extinct genus of dissorophoid temnospondyl within the family Micropholidae. It is known from the early Permian of North America.
Biseridens is an extinct genus of anomodont therapsid, and one of the most basal anomodont genera known. Originally known from a partial skull misidentified as an eotitanosuchian in 1997, another well-preserved skull was found in the Xidagou Formation, an outcropping in the Qilian Mountains of Gansu, China, in 2009 that clarified its relationships to anomodonts, such as the dicynodonts.
Raranimus is an extinct genus of therapsids of the Middle Permian. It was described in 2009 from a partial skull found in 1998 from the Dashankou locality of the Xidagou Formation, outcropping in the Qilian Mountains of Gansu, China. The genus is the most basal known member of the clade Therapsida, to which the later Mammalia belong.
Sinophoneus is an extinct genus of carnivorous dinocephalian therapsid belonging to the family Anteosauridae. It lived 272 to 270 million years ago at the beginning of the Middle Permian in what is now the Gansu Province in northern China. It is known by a skull of an adult individual, as well as by many skulls of juvenile specimens. The latter were first considered as belonging to a different animal, named Stenocybus, before being reinterpreted as immature Sinophoneus. Sinophoneus shows a combination of characters present in other anteosaurs. Its bulbous profile snout and external nostrils located in front of the canine are reminiscent of the basal anteosaur Archaeosyodon, while its massive transerse pterygoids processes with enlarged distal ends are more similar to the more derived anteosaurs Anteosaurus and Titanophoneus. First phylogenetic analyzes identified Sinophoneus as the most basal Anteosaurinae. A more recent analysis positioned it outside the Anteosaurinae and Syodontinae subclades, and recovers it as the most basal Anteosauridae.
Bunostegos is an extinct genus of pareiasaur parareptile from the Late Permian of the Agadez Region in Niger. The type species, Bunostegos akokanensis, was named from the Moradi Formation in 2003. It was a cow-sized animal with a distinctive skull that had large bony knobs, similar in form to those of other pareiasaurs but far larger. The species appears to have lived in a desert in the centre of the supercontinent of Pangaea.
Scapanops is an extinct genus of dissorophid temnospondyl amphibian known from the Early Permian Nocona Formation of north-central Texas, United States. It contains only the type species Scapanops neglecta, which was named by Rainer R. Schoch and Hans-Dieter Sues in 2013. Scapanops differs from other dissorophids in having a very small skull table, which means that its eye sockets are unusually close to the back of the skull. The eye sockets are also very large and spaced far apart. Scapanops was probably small-bodied with a proportionally large head and short trunk and tail. Like other dissorophids, it probably spent most of its life on land.
Eucacopinae is an extinct clade of dissorophid temnospondyls. Eucacopines differ from the other main group of dissorophids, the Dissorophinae, in having more lightly built skeletons and more knobby skulls. The subfamily was originally named Cacopinae, but since the name was already established for a group of living microhylid frogs in 1931, the name was changed to Eucacopinae in 2013. Eucacopinae is a stem-based taxon defined as the most inclusive clade containing the species Cacops apsidephorus but not Dissorophus multicinctus, which belongs to Dissorophinae. According to the most recent phylogenetic analyses of Dissorophidae, Eucacopinae includes the basal ("primitive") species Conjunctio multidens and Scapanops neglecta from the southwestern United States and a more derived ("advanced") group including several species of Cacops and the Russian genera Kamacops and Zygosaurus. Derived eucacopines have two rows of bony plates called osteoderms running down their backs, while the more basal eucacopines have only a single row. Dissorophines also have a double row of osteoderms but probably evolved them independently because the most recent common ancestor of the two groups had a single row of osteoderms.
Reiszerpeton is an extinct genus of dissorophid temnospondyl known from the Early Permian Archer City Formation of Texas. It is known solely from the holotype, MCZ 1911, a complete skull. This specimen was originally referred to the amphibamiform Tersomius texensis. A reappraisal of the holotype of T. texensis and a number of other referred specimens by Maddin et al. (2013) noted a number of differences from both T. texensis and amphibamiforms more broadly that suggested affinities with the Dissorophidae. This was confirmed by a phylogenetic analysis, which placed it as the sister taxon to the Eucacopinae. Reiszerpeton is known only from the type species, R. renascentis, which was named for Canadian paleontologist Robert Reisz. The species name refers to the recognition of Reisz as a "renaissance paleontologist." It is differentiated from other dissorophids by its small size, small and more numerous maxillary teeth, smooth cranial ornamentation, and greater distance between the orbit and the otic notch.
Alveusdectes is an extinct genus of diadectid tetrapod from the Late Permian of China. Like other diadectids, it was a large-bodied terrestrial herbivore capable of eating tough fibrous plant material. It was described in 2015 on the basis of a single partial skull and lower jaw found in the Shangshihezi Formation near the city of Jiyuan in Henan. This skull was found in a layer of the Shangshihezi Formation that dates to about 256 million years ago and contains the remains of many other terrestrial tetrapods including pareiasaurs, chroniosuchians, and therapsids. Alveusdectes is the youngest known diadectid by 16 million years and is also the only diadectid known from Asia. It likely represents a late-surviving lineage of diadectids that radiated eastward from western Laurasia into north China. Diadectids are otherwise absent from eastern Laurasia, which may reflect their low diversity at the time. Diadectids first appeared in the Late Carboniferous and were the first animals to have ever occupied the niche of large-bodied herbivores, allowing them to undergo an evolutionary radiation in the Early Permian. By the Late Permian many other groups of tetrapods had entered that niche, and increased competition among herbivores likely resulted in the eventual extinction of diadectids. Alveusdectes may have been able to persist because the fauna of north China seems to have been isolated from other Laurasian faunas during the Late Permian, meaning that fewer herbivores were competing for the same ecological space.
This list of fossil amphibians described in 2018 is a list of new taxa of fossil amphibians that were described during the year 2018, as well as other significant discoveries and events related to amphibian paleontology that occurred in 2018.