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Cacops, is a genus of dissorophid temnospondyls from the Kungurian stage of the early Permian of the United States. Cacops is one of the few olsoniforms whose ontogeny is known. Cacops fossils were almost exclusively known from the Cacops Bone Bed of the Lower Permian Arroyo Formation of Texas for much of the 20th century. New material collected from the Dolese Brothers Quarry, near Richards Spur, Oklahoma in the past few decades has been recovered, painting a clearer picture of what the animal looked and acted like.
Dissorophidae is an extinct family of medium-sized, temnospondyl amphibians that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America.
Temnospondyli is a diverse order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian, and Triassic periods. A few species continued into the Jurassic and Cretaceous periods. Fossils have been found on every continent. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are considered amphibians, many had characteristics, such as scales and armour-like bony plates, that distinguish them from modern amphibians (lissamphibians).
Acheloma is an extinct genus of temnospondyl that lived during the Early Permian. The type species is A. cumminsi.
Trematopidae is a family of dissorophoid temnospondyl spanning the late Carboniferous to the early Permian. Together with Dissorophidae, the family forms Olsoniformes, a clade comprising the medium-large terrestrial dissorophoids. Trematopids are known from numerous localities in North America, primarily in New Mexico, Oklahoma, and Texas, and from the Bromacker quarry in Germany.
Broiliellus is an extinct genus of dissorophoid temnospondyl within the family Dissorophidae. Broiliellus is most closely related to the genus Dissorophus, and both have been placed in the subfamily Dissorophinae. Broiliellus is known from five species from the Early Permian: the type species is Broiliellus texensis, and the other species are Broiliellus brevis,Broiliellus olsoni, Broiliellus arroyoensis, and Broiliellus reiszi. An additional species, Broiliellus novomexicanus, which was originally named Aspidosaurus novomexicanus, is now thought to fall outside the genus as a member of the subfamily Eucacopinae.
Tersomius is an extinct genus of dissorophoid temnospondyl within the family Micropholidae. It is known from the early Permian of North America.
Micropholis is an extinct genus of dissorophoid temnospondyl. Fossils have been found from the Lystrosaurus Assemblage Zone of the Karoo Basin in South Africa and are dated to the Induan. Fossils have also been found from the lower Fremouw of Antarctica.Micropholis is the only post-Permian dissorophoid and the only dissorophoid in what is presently the southern hemisphere and what would have been termed Gondwana during the amalgamation of Pangea.
Acleistorhinidae is an extinct family of Late Carboniferous and Early Permian-aged parareptiles. Acleistorhinids are most diverse from the Richards Spur locality of the Early Permian of Oklahoma. Richards Spur acleistorhinids include Acleistorhinus, Colobomycter, and possibly Delorhynchus and Feeserpeton. Other taxa include Carbonodraco from the Late Carboniferous of Ohio and Karutia from the Early Permian of Brazil. Acleistorhinidae is commonly considered a subgroup of lanthanosuchoids, related to taxa such as Chalcosaurus, Lanthaniscus and Lanthanosuchus. However, a re-examination of parareptile phylogeny conducted by Cisneros et al. (2021) argued that lanthanosuchids were not closely related to acleistorhinids. The phylogenetic analysis conducted by these authors recovered acleistorhinids as the sister group of the clade Procolophonia, while lanthanosuchids were recovered within the procolophonian subgroup Pareiasauromorpha.
The Amphibamidae are an extinct family of dissorophoid temnospondyls known from Late Carboniferous-Early Permian strata in the United States.
Rubeostratilia is an extinct genus of amphibamiform temnospondyl from the early Permian of Texas. It is known from a single skull. This genus was named by Hélène Bourget and Jason S. Anderson in 2011, and the type species is Rubeostratilia texensis. The genus name comes from the Latin translation of 'redbeds' in reference to the Texas redbeds that produced both the holotype and many other early Permian fossils. The specific name is for the state of Texas. The holotype and only known specimen was collected in 1941 from the Nocona Formation exposures in Clay County by a Works Projects Administration project that was transferred to the Field Museum of Natural History through an interinstitutional exchange with the Texas Memorial Museum.
Georgenthalia is an extinct genus of dissorophoid temnospondyl from the Lower Permian. It is an amphibamid which lived in what is now the Thuringian Forest of central Germany. It is known from the holotype MNG 11135, a small, complete skull. It was found in the Bromacker locality of the Tambach Formation. It was first named by Jason S. Anderson, Amy C. Henrici, Stuart S. Sumida, Thomas Martens and David S. Berman in 2008 and the type species is Georgenthalia clavinasica.
Plemmyradytes is an extinct genus of dissorophoid temnospondyl from the early Permian. It is an amphibamiform from the Eskridge Formation exposures of Nebraska. The type species is Plemmyradytes shintoni. The genus name derives from the Greek plemmyris and dytes ('diver'), while the specific name honors John Shinton, a fossil preparator at the Denver Museum of Natural History where all known specimens of this taxon are reposited following collection in the late 20th century.
Olsoniformes is a clade of dissorophoid temnospondyls. It includes the families Dissorophidae and Trematopidae. Most members of the clade were highly adapted to a terrestrial lifestyle. The clade was named in 2008 and is defined as the least inclusive clade containing Dissorophus multicinctus and Acheloma cumminsi but not Amphibamus grandiceps, Micromelerpeton credneri, and Apateon pedestris. Olsoniforms share various features such as a stout and low ilium and a thin cultriform process.
Perryella is an extinct genus of dvinosaurian(?) temnospondyl from the Permian of Oklahoma.
Reiszerpeton is an extinct genus of dissorophid temnospondyl known from the Early Permian Archer City Formation of Texas. It is known solely from the holotype, MCZ 1911, a complete skull. This specimen was originally referred to the amphibamiform Tersomius texensis. A reappraisal of the holotype of T. texensis and a number of other referred specimens by Maddin et al. (2013) noted a number of differences from both T. texensis and amphibamiforms more broadly that suggested affinities with the Dissorophidae. This was confirmed by a phylogenetic analysis, which placed it as the sister taxon to the Eucacopinae. Reiszerpeton is known only from the type species, R. renascentis, which was named for Canadian paleontologist Robert Reisz. The species name refers to the recognition of Reisz as a "renaissance paleontologist." It is differentiated from other dissorophids by its small size, small and more numerous maxillary teeth, smooth cranial ornamentation, and greater distance between the orbit and the otic notch.
The Micropholidae are an extinct family of dissorophoid euskelian temnospondyls known from Late Carboniferous to Early Triassic strata in the United States and South Africa.
Amphibamiformes is an unranked clade with Dissorophoidea created by Schoch (2018). It encompasses all of the taxa traditionally considered to be "amphibamids", branchiosaurids, and hypothetically lissamphibians under the traditional temnospondyl hypothesis of lissamphibian origins. These taxa are typically small-bodied dissorophoids and form the sister group to Olsoniformes, which comprises dissorophids and trematopids.
Richards Spur is a Permian fossil locality located at the Dolese Brothers Limestone Quarry north of Lawton, Oklahoma. The locality preserves clay and mudstone fissure fills of a karst system eroded out of Ordovician limestone and dolomite, with the infilling dating to the Artinskian stage of the early Permian (Cisuralian), around 289 to 286 million years ago. Fossils of terrestrial animals are abundant and well-preserved, representing one of the most diverse Paleozoic tetrapod communities known. A common historical name for the site is Fort Sill, in reference to the nearby military base. Fossils were first reported at the quarry by workers in 1932, spurring a wave of collecting by local and international geologists. Early taxa of interest included the abundant reptile Captorhinus and microsaurs such as Cardiocephalus and Euryodus. Later notable discoveries include Doleserpeton, the most diverse assortment of parareptiles in the Early Permian, and the rare early diapsid Orovenator.
Nadia Belinda Fröbisch is a German vertebrate paleontologist and developmental biologist who specializes in the evolution and development of amphibians. She is currently a professor at the Museum für Naturkunde Berlin in the Leibniz Institute for Research on Evolution and Biodiversity.
References
- ↑ Fröbisch, N.B.; Reisz, R.R. (2008). "A new Lower Permian amphibamid (Dissorophoidea, Temnospondyli) from the fissure fill deposits near Richards Spur, Oklahoma". Journal of Vertebrate Paleontology. 28 (4): 1015–1030. doi:10.1671/0272-4634-28.4.1015.
- ↑ Anderson, Jason S.; Bolt, John R. (2013). "New information on amphibamids (Tetrapoda, Temnospondyli) from Richards Spur (Fort Sill), Oklahoma". Journal of Vertebrate Paleontology. 33 (3): 553–567. doi:10.1080/02724634.2013.726676. ISSN 0272-4634.
- ↑ Maddin, Hillary C.; Fröbisch, Nadia B.; Evans, David C.; Milner, Andrew R. (2013). "Reappraisal of the Early Permian amphibamid Tersomius texensis and some referred material". Comptes Rendus Palevol. 12 (7–8): 447–461. doi:10.1016/j.crpv.2013.06.007.
- ↑ Anderson, J.S.; Reisz, R.R.; Scott, D.; Fröbisch, N.B.; Sumida, S.S. (2008). "A stem batrachian from the Early Permian of Texas and the origin of frogs and salamanders" (PDF). Nature. 453 (7194): 515–518. doi:10.1038/nature06865. PMID 18497824. Archived from the original (PDF) on 2011-07-26.
- ↑ Schoch, Rainer R. (2019). "The putative lissamphibian stem-group: phylogeny and evolution of the dissorophoid temnospondyls". Journal of Paleontology. 93 (1): 137–156. doi: 10.1017/jpa.2018.67 . ISSN 0022-3360.
