| Rotaryus Temporal range: Early Permian, | |
|---|---|
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| Restoration of Rotaryus gothae | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Order: | † Temnospondyli |
| Family: | † Trematopidae |
| Genus: | † Rotaryus Berman et al., 2011 |
| Type species | |
| †Rotaryus gothae Berman et al., 2011 | |
Rotaryus is an extinct genus of Early Permian trematopid dissorophoid temnospondyl known from the Free State of Thuringia of central Germany. [1]
Rotaryus is known only from the holotype MNG 10182, articulated partial well-preserved skull and both mandibles and a closely associated partial postcranial skeleton. The postcranial skeleton includes several articulated neural arches with ribs, most of the left shoulder girdle, humeri, right radius and ulna, and a femur. It was collected from the uppermost part of the Tambach Formation, dating to the Artinskian stage of the Late Cisuralian Series (or alternatively upper Rotliegend), about 284-279.5 million years ago. It was found in the lowermost formational unit of the Upper Rotliegend Group or Series of the Bromacker Quarry, the middle part of the Thuringian Forest, near the village of Tambach-Dietharz. Rotaryus is only the second trematopid species to be reported from the Bromacker locality, and outside of United States, alongside Tambachia . [1]
Rotaryus is characterized by a unique combination of characters, including the two following autapomorphies: "entire length of nasolacrimal canal exposed as a smooth, uniform channel in which the anterior half coincides with the maxillary-lacrimal suture; and parasphenoid plate has an outline of an isosceles triangle, with the shorter, equal-length lateral margins converging on the cultriform process." The following cladogram shows the phylogenetic position of Rotaryus, from Berman et al., 2011. [1]
| Dissorophoidea |
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Rotaryus was first named by David S. Berman, Amy C. Henrici, Thomas Martens, Stuart S. Sumida and Jason S. Anderson in 2011 and the type species is Rotaryus gothae. Both the generic name and the specific name honor the Rotary Club of Gotha, Germany, for supporting financially the excavations and the Bromacker locality project. [1]
Diadectidae is an extinct family of early tetrapods that lived in what is now North America and Europe during the Late Carboniferous and Early Permian, and in Asia during the Late Permian. They were the first herbivorous tetrapods, and also the first fully terrestrial animals to attain large sizes. Footprints indicate that diadectids walked with an erect posture. They were the first to exploit plant material in terrestrial food chains, making their appearance an important stage in both vertebrate evolution and the development of terrestrial ecosystems.
Seymouria is an extinct genus of Seymouriamorph from the Early Permian of North America and Europe. Although they were amphibians, Seymouria were well-adapted to life on land, with many reptilian features—so many, in fact, that Seymouria was first thought to be a primitive reptile. It is primarily known from two species, Seymouria baylorensis and Seymouria sanjuanensis. The type species, S. baylorensis, is more robust and specialized, though its fossils have only been found in Texas. On the other hand, Seymouria sanjuanensis is more abundant and widespread. This smaller species is known from multiple well-preserved fossils, including a block of six skeletons found in the Cutler Formation of New Mexico, and a pair of fully grown skeletons from the Tambach Formation of Germany, which were fossilized lying next to each other.
Caseasauria is one of the two main clades of early synapsids, the other being the Eupelycosauria. Caseasaurs are currently known only from the Late Carboniferous and the Permian, and include two superficially different families, the small insectivorous or carnivorous Eothyrididae, and the large, herbivorous Caseidae. These two groups share a number of specialised features associated with the morphology of the snout and external naris.
Eudibamus is an extinct genus of biped bolosaurid ankyramorph parareptile known from the Free State of Thuringia of central Germany. It had a very small size reaching only 25-26 cm in length.
Orobates is an extinct genus of diadectid. It lived in the middle Permian, about 260 million years ago. Its remains were found in Germany. A combination of primitive and derived traits distinguish it from all other well-known members of the Diadectidae, a family of herbivorous reptile-like amphibians. It weighed about 4 kg and appears to have been part of an upland fauna, browsing on high fibre plants.
Trematopidae is a family of dissorophoid temnospondyl spanning the late Carboniferous to the early Permian. Together with Dissorophidae, the family forms Olsoniformes, a clade comprising the medium-large terrestrial dissorophoids. Trematopids are known from numerous localities in North America, primarily in New Mexico, Oklahoma, and Texas, and from the Bromacker quarry in Germany.
Anconastes is an extinct genus of dissorophoid temnospondyl within the family Trematopidae. It is known from two specimens from the Late Carboniferous Cutler Formation of north-central New Mexico in the southwestern United States. The genus name derives from two Greek roots, ankos and nastes ("inhabitant"), which refers to the type locality of El Cobre Canyon where the specimens were found. The species name derives from the Latin word vesperus ("western"). The more complete specimen, the holotype, is a partial skull with articulated mandibles and a substantial amount of the postcranial skeleton. The less complete specimen, the paratype, consists only of the right margin of the skull with an articulated mandible.
Actiobates is an extinct genus of trematopid temnospondyl that lived during the Late Carboniferous. It is known from the Garnett Quarry in Kansas.
Ecolsonia is an extinct genus of trematopid temnospondyl. Its phylogenetic position within Olsoniformes has been historically debated, but it is presently considered to be a trematopid.
Tambachia is an extinct genus of dissorophoid temnospondyl within the family Trematopidae. It is known from the Early Permian Tambach Formation near the town of Tambach-Dietharz in Thuringia, Germany. Tambachia is the first trematopid to have been discovered outside the United States.
Ostodolepidae, also spelled Ostodolepididae, is an extinct family of Early Permian microsaurs. They are unique among microsaurs in that they were large, reaching lengths of up to 2 feet (61 cm), terrestrial, and presumably fossorial. Ostodolepid remains have been found from Early Permian beds in Texas, Oklahoma, and Germany.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2011.
Georgenthalia is an extinct genus of dissorophoid temnospondyl from the Lower Permian. It is an amphibamid which lived in what is now the Thuringian Forest of central Germany. It is known from the holotype MNG 11135, a small, complete skull. It was found in the Bromacker locality of the Tambach Formation. It was first named by Jason S. Anderson, Amy C. Henrici, Stuart S. Sumida, Thomas Martens and David S. Berman in 2008 and the type species is Georgenthalia clavinasica.
Olsoniformes is a clade of dissorophoid temnospondyls. It includes the families Dissorophidae and Trematopidae. Most members of the clade were highly adapted to a terrestrial lifestyle. The clade was named in 2008 and is defined as the least inclusive clade containing Dissorophus multicinctus and Acheloma cumminsi but not Amphibamus grandiceps, Micromelerpeton credneri, and Apateon pedestris. Olsoniforms share various features such as a stout and low ilium and a thin cultriform process.
Tambaroter is an extinct genus of ostodolepid microsaur from the Early Permian of Germany. The type species T. carrolli was named in 2011. Tambaroter is known from a single skull found in the Tambach Formation, which is the lowermost unit of the Upper Rotliegend. It is the only vertebrate that has been found outside the Bromacker Quarry, the most productive locality of the formation. It is also the first ostodolepid known from outside North America.
Thuringothyris is an extinct genus of Early Permian eureptiles known from the Thuringian Forest in central Germany.
The Tambach Formation is an Early Permian-age geologic formation in central Germany. It consists of red to brown-colored sedimentary rocks such as conglomerate, sandstone, and mudstone, and is the oldest portion of the Upper Rotliegend within the Thuringian Forest Basin.
Recumbirostra is a clade of tetrapods which lived during the Carboniferous and Permian periods. It includes the families Pantylidae, Gymnarthridae, Ostodolepidae, Rhynchonkidae and Brachystelechidae, with additional families such as Microbrachidae and Molgophidae being included by some authors. Recumbirostra was erected as a clade in 2007 to include many of the taxa traditionally grouped in "Microsauria", which has since been shown to be a paraphyletic or polyphyletic grouping. Like other "microsaurs", the recumbirostrans have traditionally been considered to be members of the subclass Lepospondyli; however, phylogenetic analyses conducted by Pardo, Szostakiwskyj and Anderson (2015) and Pardo et al. (2017) recovered them as early-branching sauropsid amniotes instead.
Eucacopinae is an extinct clade of dissorophid temnospondyls. Eucacopines differ from the other main group of dissorophids, the Dissorophinae, in having more lightly built skeletons and more knobby skulls. The subfamily was originally named Cacopinae, but since the name was already established for a group of living microhylid frogs in 1931, the name was changed to Eucacopinae in 2013. Eucacopinae is a stem-based taxon defined as the most inclusive clade containing the species Cacops apsidephorus but not Dissorophus multicinctus, which belongs to Dissorophinae. According to the most recent phylogenetic analyses of Dissorophidae, Eucacopinae includes the basal ("primitive") species Conjunctio multidens and Scapanops neglecta from the southwestern United States and a more derived ("advanced") group including several species of Cacops and the Russian genera Kamacops and Zygosaurus. Derived eucacopines have two rows of bony plates called osteoderms running down their backs, while the more basal eucacopines have only a single row. Dissorophines also have a double row of osteoderms but probably evolved them independently because the most recent common ancestor of the two groups had a single row of osteoderms.
Tambacarnifex is an extinct genus of varanodontine synapsids known from the Early Permian Tambach Formation of Free State of Thuringia, central Germany. It was first named by David S. Berman, Amy C. Henrici, Stuart S. Sumida, Thomas Martens and Valerie Pelletier in 2013 and the type species is Tambacarnifex unguifalcatus.
