| Ardenna | |
|---|---|
 | |
| Great shearwater (Ardenna gravis) | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Aves |
| Order: | Procellariiformes |
| Family: | Procellariidae |
| Genus: | Ardenna Reichenbach, 1853 |
| Type species | |
| Puffinus major Faber, 1822 [1] = Procellaria gravis O'Reilly, 1818 | |
| Species | |
See text | |
Ardenna is a genus of seabirds in the family Procellariidae. These medium-sized to large shearwater species were formerly included in the genus Puffinus .
A phylogenetic analysis using mitochondrial DNA published in 2004 found that Puffinus contained two distinct clades and was polyphyletic, with Puffinus more closely related to Calonectris than to Ardenna. [2] [3] To create monophyletic genera a group of species were moved into Ardenna, a genus that had been introduced in 1853 by Ludwig Reichenbach with the great shearwater as the type species. [4] [5] Reichenbach cites the Italian naturalist Ulisse Aldrovandi who in 1603 used the spelling "Artenna" for a seabird. [6] [7] Recent genomic studies have validated the phylogenetic distinction between Ardenna and Puffinus, though this study finding Ardenna rather than Puffinus being closer to Calonectris. [8]
The genus contains seven extant species as shown below: [9]
| Image | Scientific name | Common Name | Distribution [10] |
|---|---|---|---|
 | A. pacifica | Wedge-tailed shearwater | Breeds on islands in warm temperate and tropical areas of the Indian and Pacific Oceans between 30°N and 30°S, foraging in surrounding waters. |
 | A. bulleri | Buller's shearwater | Breeds New Zealand, foraging across most of the Pacific Ocean north to 60°N. |
 | A. grisea | Sooty shearwater | Breeds Tierra del Fuego, Falkland Islands, southeast Australia, New Zealand, foraging widespread in Atlantic and Pacific Oceans north to 65°N (rarely 70°N). |
 | A. tenuirostris | Short-tailed shearwater | Breeds southern Australia, foraging across most of the western and central Pacific Ocean and marginally into the Arctic Ocean, north to 70°N. |
 | A. creatopus | Pink-footed shearwater | Breeds Juan Fernandez and Mocha Islands off Chile, foraging in the eastern Pacific Ocean north to 60°N. |
 | A. carneipes | Flesh-footed shearwater | Breeds southern Indian Ocean (Amsterdam Island) and southwest Pacific Ocean including Lord Howe Island, South Australia and northern New Zealand, foraging across most of the western and central Pacific Ocean north to 60°N. |
 | A. gravis | Great shearwater | Breeds southern Atlantic Ocean on Tristan da Cunha group, also small numbers on the Falkland Islands, foraging across most of the Atlantic Ocean north to 65°N. |
The genus contains five extinct species as shown below: [11]
| Scientific name | Stratigraphic age | Distribution |
|---|---|---|
| A. buchananbrowni | Late Pliocene | New Zealand |
| A. conradi | Middle Miocene | Maryland, USA |
| A. davealleni | Late Pliocene | New Zealand |
| A. gilmorei | Late Pliocene | California, USA |
| A. pacificoides | Pleistocene | Saint Helena |
Phylogeny based on a study by Joan Ferrer Obiol and collaborators published in 2022. [12]
| Ardenna |
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Shearwaters are medium-sized long-winged seabirds in the petrel family Procellariidae. They have a global marine distribution, but are most common in temperate and cold waters, and are pelagic outside the breeding season.
Petrels are tube-nosed seabirds in the phylogenetic order Procellariiformes.
The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prions, and the shearwaters. This family is part of the bird order Procellariiformes, which also includes the albatrosses and the storm petrels.
The Balearic shearwater is a medium-sized shearwater in the seabird family Procellariidae. Puffinus is a Neo-Latin loanword based on the English "puffin" and its variants, that referred to the cured carcass of the fat nestling of the Manx shearwater, a former delicacy. The specific mauretanicus refers to Mauretania, an old name for an area of North Africa roughly corresponding to Morocco and Algeria. The Balearic Shearwater is listed critically endangered by the IUCN and is one of Europe's most endangered seabirds.
The great shearwater is a large shearwater in the seabird family Procellariidae. It breeds colonially on rocky islands in the south Atlantic. Outside the breeding season it ranges widely in the Atlantic.
Cory's shearwater is a large shearwater in the seabird family Procellariidae. It breeds colonially on the archipelago of the Azores in the eastern Atlantic. Outside the breeding season it ranges widely in the Atlantic. It was formerly considered to be conspecific with Scopoli's shearwater.
The sooty shearwater is a medium-large shearwater in the seabird family Procellariidae. In New Zealand, it is also known by its Māori name tītī, and is harvested by Māori people for muttonbird, like its relatives the wedge-tailed shearwater and the Australian short-tailed shearwater.
The little shearwater is a small shearwater in the petrel family Procellariidae. Despite the generic name, it is unrelated to the puffins, which are auks, the only similarity being that they are both burrow-nesting seabirds.
The yelkouan shearwater, Levantine shearwater or Mediterranean shearwater is a medium-sized shearwater in the seabird family Procellariidae. It was formerly treated as a subspecies of the Manx shearwater.
Puffinus is a genus of seabirds in the order Procellariiformes that contains about 20 small to medium-sized shearwaters. Two other shearwater genera are named: Calonectris, which comprises three or four large shearwaters, and Ardenna with another seven species.
Calonectris is a genus of seabirds. The genus name comes from Ancient Greek kalos, "good" and nectris, "swimmer".
The wedge-tailed shearwater is a medium-large shearwater in the seabird family Procellariidae. It is one of the shearwater species that is sometimes referred to as a muttonbird, like the sooty shearwater of New Zealand and the short-tailed shearwater of Australia. It is found throughout the tropical Pacific and Indian Oceans, roughly between latitudes 35°N and 35°S. It breeds on the islands off Japan, on the Islas Revillagigedo, the Hawaiian Islands, the Seychelles, the Northern Mariana Islands, and off Eastern and Western Australia.
The short-tailed shearwater or slender-billed shearwater, also called yolla or moonbird, and commonly known as the muttonbird in Australia, is the most abundant seabird species in Australian waters, and is one of the few Australian native birds in which the chicks are commercially harvested. It is a migratory species that breeds mainly on small islands in Bass Strait and Tasmania and migrates to the Northern Hemisphere for the boreal summer.
Townsend's shearwater is a rare seabird of the tropics from the family Procellariidae.
Procellaria is a genus of Southern Ocean long-winged seabirds related to prions, and within the order Procellariiformes. The black petrel ranges in the Pacific Ocean, and as far north as Central America. The spectacled petrel is confined to the Atlantic Ocean, and the Westland petrel to the Pacific Ocean. The white-chinned and grey petrel range throughout the higher latitudes of the Southern Ocean.
The flesh-footed shearwater is a medium-sized shearwater. Its plumage is black. It has pale pinkish feet, and a pale bill with a distinct black tip. Together with the equally light-billed pink-footed shearwater, it forms the Hemipuffinus group, a superspecies which may or may not have an Atlantic relative in the great shearwater. These large shearwaters are among those that have been separated into the genus Ardenna. Recent genetic analysis indicates evidence of strong divergence between Pacific colonies relative to those in South and Western Australia, thought to be explained by philopatry and differences in foraging strategies during the breeding season.
Buller's shearwater is a Pacific species of seabird in the family Procellariidae; it is also known as the grey-backed shearwater or New Zealand shearwater. A member of the black-billed wedge-tailed Thyellodroma group, among the larger shearwaters of the genus Ardenna, it forms a superspecies with the wedge-tailed shearwater.
The pink-footed shearwater is a species of seabird. The bird is 48 cm (19 in) in length, with a 109 cm (43 in) wingspan. It is polymorphic, having both darker- and lighter-phase populations. Together with the equally light-billed flesh-footed shearwater, it forms the Hemipuffinus group, a superspecies that may or may not have an Atlantic relative in the great shearwater. These are large shearwaters which are among those that could be separated in the genus Ardenna.
The Rapa shearwater, is a rare seabird of the tropics from the family Procellariidae. It breeds on the surrounding islets of Rapa in the Austral Islands of French Polynesia where it is known locally as the kaki kaki.
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