Camelini

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Camelini
Temporal range: Miocene–Recent
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Camelus dromedarius
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Family: Camelidae
Subfamily: Camelinae
Tribe: Camelini
Gray, 1821 [1]
Genera

Camelini is a tribe of camelids including all camelids more closely related to modern camels (Camelus) than to Lamini (which contains llamas, alpacas, vicuñas, and guanacos), from which camelines split at least 16 million years ago. [2] [3] The tribe originated in North America, with the genus Paracamelus migrating over the Bering Land Bridge into Eurasia during the Late Miocene, around 6 million years ago, becoming ancestral to Camelus. [4] [5] [6] The last member of Camelini in North America was Camelops , which became extinct as part of the Quaternary extinction event at the end of the Late Pleistocene, around 12,000 years ago. [3]

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<span class="mw-page-title-main">Camel</span> Genus of mammals

A camel is an even-toed ungulate in the genus Camelus that bears distinctive fatty deposits known as "humps" on its back. Camels have long been domesticated and, as livestock, they provide food and textiles. Camels are working animals especially suited to their desert habitat and are a vital means of transport for passengers and cargo. There are three surviving species of camel. The one-humped dromedary makes up 94% of the world's camel population, and the two-humped Bactrian camel makes up 6%. The wild Bactrian camel is a separate species and is now critically endangered.

<span class="mw-page-title-main">Artiodactyl</span> Order of mammals

Artiodactyls are placental mammals belonging to the order Artiodactyla. Typically, they are ungulates which bear weight equally on two of their five toes: the third and fourth, often in the form of a hoof. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, most perissodactyls bear weight on an odd number of the five toes. Another difference between the two is that many artiodactyls digest plant cellulose in one or more stomach chambers rather than in their intestine as perissodactyls do. The advent of molecular biology, along with new fossil discoveries, found that cetaceans fall within this taxonomic branch, being most closely related to hippopotamuses. Some modern taxonomists thus apply the name Cetartiodactyla to this group, while others opt to include cetaceans within the existing name of Artiodactyla. Some researchers use "even-toed ungulates" to exclude cetaceans and only include terrestrial artiodactyls, making the term paraphyletic in nature.

<i>Smilodon</i> Extinct genus of saber-toothed cat

Smilodon is a genus of felids belonging to the extinct subfamily Machairodontinae. It is one of the best known saber-toothed predators and prehistoric mammals. Although commonly known as the saber-toothed tiger, it was not closely related to the tiger or other modern cats. Smilodon lived in the Americas during the Pleistocene epoch. The genus was named in 1842 based on fossils from Brazil; the generic name means "scalpel" or "two-edged knife" combined with "tooth". Three species are recognized today: S. gracilis, S. fatalis, and S. populator. The two latter species were probably descended from S. gracilis, which itself probably evolved from Megantereon. The hundreds of specimens obtained from the La Brea Tar Pits in Los Angeles constitute the largest collection of Smilodon fossils.

<span class="mw-page-title-main">Tylopoda</span> Suborder of mammals

Tylopoda is a suborder of terrestrial herbivorous even-toed ungulates belonging to the order Artiodactyla. They are found in the wild in their native ranges of South America and Asia, while Australian feral camels are introduced. The group has a long fossil history in North America and Eurasia. Tylopoda appeared during the Eocene around 50 million years ago.

<span class="mw-page-title-main">Camelidae</span> Family of mammals

Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, and guanacos. Camelids are even-toed ungulates classified in the order Artiodactyla, along with species including whales, pigs, deer, cattle, and antelopes.

<span class="mw-page-title-main">Bactrian camel</span> Central Asian mammal, beast of burden

The Bactrian camel, also known as the Mongolian camel, domestic Bactrian camel or two-humped camel, is a large even-toed ungulate native to the steppes of Central Asia. It has two humps on its back, in contrast to the single-humped dromedary. Its population of 2 million exists mainly in the domesticated form. Their name comes from the ancient historical region of Bactria.

<span class="mw-page-title-main">Gomphothere</span> Extinct family of proboscidean mammals

Gomphotheres are an extinct group of proboscideans related to modern elephants. They were widespread across Afro-Eurasia and North America during the Miocene and Pliocene epochs and dispersed into South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group that is ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae. While most famous forms such as Gomphotherium had long lower jaws with tusks, which is the ancestral condition for the group, some later members developed shortened (brevirostrine) lower jaws with either vestigial or no lower tusks, looking very similar to modern elephants, an example of parallel evolution, which outlasted the long-jawed gomphotheres. By the end of the Early Pleistocene, gomphotheres became extinct in Afro-Eurasia, with the last two genera, Cuvieronius ranging from southern North America to western South America, and Notiomastodon having a wide range over most of South America until the end of the Pleistocene around 12,000 years ago, when they became extinct following the arrival of humans.

<i>Tomistoma</i> Genus of crocodilians

Tomistoma is a genus of gavialid crocodilians. They are noted for their long narrow snouts used to catch fish, similar to the gharial. Tomistoma contains one extant (living) member, the false gharial, as well as potentially several extinct species: T. cairense, T. lusitanicumT. coppensi, and T. dowsoni. However, these species may need to be reclassified to different genera as studies have shown them to be paraphyletic, for example: previously assigned species T. taiwanicus from Taiwan, is reclassified to the genus Toyotamaphimeia, and T. dowsoni should be excluded from Tomistoma based on phylogenetic analysis.

<i>Camelops</i> Extinct genus of mammals

Camelops is an extinct genus of camels that lived in North and Central America, ranging from Alaska to Honduras, from the middle Pliocene to the end of the Pleistocene. It is more closely related to the Old World dromedary and bactrian and wild bactrian camels than to the New World guanaco, vicuña, alpaca and llama; making it a true camel of the Camelini tribe. Its name is derived from the Ancient Greek κάμηλος and ὄψ, i.e. "camel-face".

<i>Titanotylopus</i> Extinct genus of mammals

Titanotylopus is an extinct genus of camel, endemic to North America from the late Hemphillian stage of the Miocene through the Irvingtonian stage of the Pleistocene. It was one of the last surviving North American camels; after its extinction, only Camelops remained.

<i>Hippidion</i> Extinct genus of mammals

Hippidion is an extinct genus of equine that lived in South America from the Late Pliocene to the end of the Late Pleistocene (Lujanian), between 2.5 million and 11,000 years ago. They were one of two lineages of equines native to South America during the Pleistocene epoch, alongside Equus (Amerhippus) neogeus.

<span class="mw-page-title-main">Caninae</span> Subfamily of carnivores

The Caninae, known as canines, are one of three subfamilies found within the canid family. The other two canid subfamilies are the extinct Borophaginae and Hesperocyoninae. The Caninae includes all living canids and their most recent fossil relatives. Their fossils were first found in North America and dated to the Oligocene era, then spreading to Asia at the end of the Miocene era, some 7 million to 8 million years ago.

<i>Bootherium</i> Extinct genus of bovid mammal

Bootherium is an extinct bovid genus from the middle to late Pleistocene of North America which contains a single species, Bootherium bombifrons. Vernacular names for Bootherium include Harlan's muskox, woodox, woodland muskox, helmeted muskox, or bonnet-headed muskox.

<span class="mw-page-title-main">Alligatoroidea</span> Superfamily of reptiles

Alligatoroidea is one of three superfamilies of crocodylians, the other two being Crocodyloidea and Gavialoidea. Alligatoroidea evolved in the Late Cretaceous period, and consists of the alligators and caimans, as well as extinct members more closely related to the alligators than the two other groups.

<i>Hemiauchenia</i> Extinct genus of mammals

Hemiauchenia is a genus of laminoid camelids that evolved in North America in the Miocene period about 10 million years ago. This genus diversified and expanded into to South America in the Late Pliocene approximately 3 to 2 million years ago, as part of the Great American Biotic Interchange. The genus became extinct at the end of the Pleistocene. The monophyly of the genus has been considered questionable, with phylogenetic analyses finding the genus to paraphyletic or polyphyletic, with some species suggested to be more closely related to living lamines than to other Hemiaucenia species.

Natalia Rybczynski is a Canadian paleobiologist, professor and researcher.

Megacamelus is an extinct genus of terrestrial herbivore in the family Camelidae, endemic to North America from the Miocene through Pliocene 10.3—4.9 mya, existing for approximately 5.4 million years.

Paracamelus is an extinct genus of camel in the family Camelidae. It originated in North America and crossed the Beringian land bridge into Eurasia during the Late Miocene, approximately 7.5–6.5 million years ago (Ma). It is the presumed ancestor to living camels of the genus Camelus.

<i>Protarctos</i> Extinct genus of bear

Protarctos is an extinct genus of basal ursine bear that lived in North America and Eurasia during the Pliocene and into Early Pleistocene.

Camelus thomasi is an extinct species of camel from the Early-Mid Pleistocene of North Africa. It is known primarily from Tighennif (Ternifine) in Algeria. Fossils from northern Sudan and Israel dated to the Late Pleistocene have been included under C. thomasi, but they are now considered to belong to different species, making C. thomasi a strictly Northwest African species.

References

  1. McKenna, Malcolm Carnegie; Bell, Susan K. (1997). Classification of Mammals: Above the Species Level. New York: Columbia University Press. p. 631. ISBN   978-0-231-11013-6.
  2. Lynch, Sinéad; Sánchez-Villagra, Marcelo R.; Balcarcel, Ana (December 2020). "Description of a fossil camelid from the Pleistocene of Argentina, and a cladistic analysis of the Camelinae". Swiss Journal of Palaeontology. 139 (1): 5. doi: 10.1186/s13358-020-00208-6 . ISSN   1664-2376. PMC   7590954 . PMID   33133011.
  3. 1 2 Buckley, Michael; Lawless, Craig; Rybczynski, Natalia (March 2019). "Collagen sequence analysis of fossil camels, Camelops and c.f. Paracamelus, from the Arctic and sub-Arctic of Plio-Pleistocene North America". Journal of Proteomics. 194: 218–225. doi:10.1016/j.jprot.2018.11.014.
  4. Heintzman, Peter D.; Zazula, Grant D.; Cahill, James A.; Reyes, Alberto V.; MacPhee, Ross D.E.; Shapiro, Beth (September 2015). "Genomic Data from Extinct North American Camelops Revise Camel Evolutionary History". Molecular Biology and Evolution. 32 (9): 2433–2440. doi:10.1093/molbev/msv128. ISSN   0737-4038. PMID   26037535.
  5. Rybczynski, Natalia; Gosse, John C.; Richard Harington, C.; Wogelius, Roy A.; Hidy, Alan J.; Buckley, Mike (June 2013). "Mid-Pliocene warm-period deposits in the High Arctic yield insight into camel evolution". Nature Communications. 4 (1): 1550. Bibcode:2013NatCo...4.1550R. doi:10.1038/ncomms2516. ISSN   2041-1723. PMC   3615376 . PMID   23462993.
  6. Singh; Tomar. Evolutionary Biology (8th revised ed.). New Delhi: Rastogi Publications. p. 334. ISBN   9788171336395.