Bactrian camel

Last updated

Bactrian camel
2011 Trampeltier 1528.JPG
Shanghai Zoo
Domesticated
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Family: Camelidae
Genus: Camelus
Species:
C. bactrianus
Binomial name
Camelus bactrianus
Camelus bactrianus distribution map.png
Synonyms [1]
List
  • Camelus bocharius Kolenati (1847)
  • Camelus caucasicus Kolenati (1847)
  • Camelus genuinus Kolenati (1847)
  • Camelus orientalis J. Fischer (1829)
  • Camelus tauricus J. Fischer (1829)

The Bactrian camel (Camelus bactrianus), also known as the Mongolian camel, domestic Bactrian camel or two-humped camel, is a large camel native to the steppes of Central Asia. It has two humps on its back, in contrast to the single-humped dromedary. [a] Its population of 2 million exists mainly in the domesticated form. [2] Their name comes from the ancient historical region of Bactria. [3]

Contents

Domesticated Bactrian camels have served as pack animals in inner Asia since ancient times. With its tolerance for cold, drought, and high altitudes, it enabled the travel of caravans on the Silk Road. Bactrian camels, whether domesticated or feral, are a separate species from the wild Bactrian camel, which is the only truly wild (as opposed to feral) species of camelid in the Old World. Domestic Bactrian camels do not descend from wild Bactrian camels, with the two species having split around 1 million years ago.

Taxonomy

Phylogenetic relationships of domestic Bactrian camels compared to living and recently extinct camels [4]
Camelidae

Lamini (llamas)

Camelini

† Camelops

Camelus

Camelus dromedarius (dromedary)

† Camelus knoblochi

Camelus ferus (wild Bactrian camel)

Camelus bactrianus (domestic Bactrian camel)

The Bactrian camel shares the genus Camelus with the dromedary (C. dromedarius) and the wild Bactrian camel (C. ferus). The Bactrian camel belongs to the family Camelidae. [1] [5] The ancient Greek philosopher Aristotle was the first European to describe the camels: in his 4th century BCE History of Animals , he identified the one-humped Arabian camel and the two-humped Bactrian camel. [6] [7] The Bactrian camel was given its current binomial name Camelus bactrianus by Swedish zoologist Carl Linnaeus in his 1758 publication Systema Naturae . [8]

Though sharing a closer common ancestor with it than with the dromedary, the domestic Bactrian camel does not descend from the wild Bactrian camel, with the two species having diverged hundreds of thousands of years ago, with their mitochondrial genomes estimated to have diverged around 1 million years ago. Genetic evidence suggests that both Bactrian camel species are closely related to the extinct giant camel species Camelus knoblochi which became extinct around 20,000 years ago, which is equidistant from both living Bactrian camel species. [4]

The Bactrian camel and the dromedary often interbreed to produce fertile offspring. Where the ranges of the two species overlap, such as in northern Punjab, Iran and Afghanistan, the phenotypic differences between them tend to decrease as a result of extensive crossbreeding between them. The fertility of their hybrid has given rise to speculation that the Bactrian camel and the dromedary should be merged into a single species with two varieties. [9] However, a 1994 analysis of the mitochondrial cytochrome b gene revealed that the species display 10.3% divergence in their sequences. [10]

Description

Camel skeleton at MAV-USP.jpg

The Bactrian camel is the largest mammal in its native range and is the largest living camel while being shorter at the shoulder than the dromedary. Shoulder height is from 160 to 180 cm (5.2 to 5.9 ft) with the overall height ranging from 230 to 250 cm (7.5 to 8.2 ft), [11] [12] [13] head-and-body length is 225–350 cm (7.38–11.48 ft), and the tail length is 35–55 cm (14–22 in). At the top of the humps, the average height is 213 cm (6.99 ft).

Bactrian camel (Camelus bactrianus).JPG

Body mass can range from 300 to 1,000 kg (660 to 2,200 lb), with males weighing around 600 kg (1,300 lb), and females around 480 kg (1,060 lb). [14] [2] [15] Its long, wooly coat varies in colour from dark brown to sandy beige. A mane and beard of long hair occurs on the neck and throat, with hairs measuring up to 25 cm (9.8 in) long.

The shaggy winter coat is shed extremely rapidly, with huge sections peeling off at once, appearing as if sloppily shorn. The two humps on the back are composed of fat (not water as is sometimes thought). The face is typical of a camelid, being long and somewhat triangular, with a split upper lip. The long eyelashes, along with the sealable nostrils, help to keep out dust in the frequent sandstorms which occur in their natural range. The two broad toes on each foot have undivided soles and are able to spread widely as an adaptation to walking on sand. The feet are very tough, as befits an animal of extreme environments.

Natural habitat

These camels are migratory, and their habitat ranges from rocky mountain massifs to flat steppe, arid desert, (mostly the Gobi Desert), stony plains and sand dunes. Conditions are extremely harsh – vegetation is sparse, water sources are limited and temperatures are extreme. The coat of the Bactrian camel can withstand cold as low as −30 °C (−22 °F) in winter to 50 °C (122 °F) in summer. [16] The camels' distribution is linked to the availability of water, with large groups congregating near rivers after rain or at the foot of the mountains, where water can be obtained from springs in the summer months, and in the form of snow during the winter.

Life history

A young calf in western Mongolia Camelus bactrianus in western Mongolia 01.jpg
A young calf in western Mongolia

Bactrian camels are exceptionally adept at withstanding wide variations in temperature, ranging from freezing cold to blistering heat. They have a remarkable ability to go without water for months at a time, but when water is available they may drink up to 57 liters at once. When well fed, the humps are plump and erect, but as resources decline, the humps shrink and lean to the side. When moving faster than a walking speed, they pace, by stepping forwards with both legs on the same side (as opposed to trotting, using alternate diagonals as done by most other quadrupeds). Speeds of up to 65 kilometres per hour (40 mph) have been recorded, but they rarely move this fast. Bactrian camels are also said to be good swimmers. The sense of sight is well developed and the sense of smell is extremely good. The lifespan of Bactrian camels is estimated at up to 50 years, more often 20 to 40 in captivity.

Diet

Skull Camel skull 2.JPG
Skull

Bactrian camels are diurnal, sleeping in the open at night and foraging for food during the day. They are primarily herbivorous. With tough mouths that can withstand sharp objects such as thorns, they are able to eat plants that are dry, prickly, salty or bitter, and can ingest virtually any kind of vegetation. When other nutrient sources are not available, these camels may feed on carcasses, gnawing on bones, skin, or various different kinds of flesh. In more extreme conditions, they may eat any material they find, which has included rope, sandals, and even tents. Their ability to feed on a wide range of foods allows them to live in areas with sparse vegetation. The first time food is swallowed, it is not fully chewed. The partly masticated food (called cud) goes into the stomach and later is brought back up for further chewing.

Bactrian camels belong to a fairly small group of animals that regularly eat snow to provide their water needs. Animals living above the snowline may have to do this, as snow and ice can be the only forms of water during winter, and by doing so, their range is greatly enlarged. The latent heat of snow and ice is big compared with the heat capacity of water, forcing animals to eat only small amounts at a time. [17]

Reproduction

Mother and calf Babybactrian.jpg
Mother and calf

Bactrian camels are induced ovulators – they ovulate after insemination (insertion of semen into the vagina); the seminal plasma, not the spermatozoa, induces ovulation. Ovulation occurs in 87% of females after insemination: 66% ovulate within 36 hours and the rest by 48 hours (the same as natural mating). The least amount of semen required to elicit ovulation is about 1.0 ml. [18]

Males during mating time are often quite violent and may bite, spit, or attempt to sit on other male camels. The age of sexual maturity varies, but is usually reached at 3–5 years. Gestation lasts around 13 months. One or occasionally two calves are produced, and the female can give birth to a new calf every other year. Young Bactrian camels are precocial, being able to stand and run shortly after birth, and are fairly large at an average birth weight of 36 kg (79 lb). They are nursed for about 1.5 years. The young calf stays with its mother for three to five years, until it reaches sexual maturity, and often helps raise subsequent generations for those years. Wild camels sometimes breed with domesticated or feral camels.

Genome

Cotswold Wildlife Park Bactrian.camel.twice.arp.jpg
Cotswold Wildlife Park

The Bactrian Camels Genome Sequencing and Analysis Consortium provides a C. bactrianus ferus genome using next generation sequencing. [19]

Several effective population size studies have been carried out. [20] They show several bottlenecks in both wild and domesticated Bactrians over the past 350,000 years. [20]

Relationship to humans

Petroglyphs Camel - panoramio (2).jpg
Petroglyphs

The Bactrian camel was domesticated circa ~4,500 BCE. [21] The dromedary is believed to have been domesticated between 4000 BCE and 2000 BCE in Arabia. As pack animals, these ungulates are virtually unsurpassed, able to carry 170–250 kg (370–550 lb) at a rate of 47 kilometres (29 mi) per day, or 4 kilometres per hour (2.5 mph) over a period of four days. [22] The species was a mainstay of transportation on the Silk Road. [23] Furthermore, Bactrian camels are frequently ridden, especially in desertified areas. In ancient Sindh, for example, Bactrian camels of two humps were initially used by the rich for riding. The camel was later brought to other areas such as Balochistan and Iran for the same purpose. [22]

Tang dynasty model, owned by Agatha Christie Greenway BactrianCamel AgathaChristie January2024 NT CCBYSA open.jpg
Tang dynasty model, owned by Agatha Christie

Bactrian camels have been the focus of artwork throughout history. For example, westerners from the Tarim Basin and elsewhere were depicted in numerous ceramic figurines of the Chinese Tang dynasty (618–907).[ citation needed ]

United States imports

Bactrian camels were imported to the U.S. several times in the mid- to late 19th century, both by the U.S. military and by merchants and miners, looking for pack animals sturdier and hardier than horses and mules. Although the camels met these needs, the United States Camel Corps was never considered much of a success. Having brought two shipments of fewer than 100 camels to the U.S., plans were made to import another 1,000, but the US Civil War interrupted this. Most surviving camels of these endeavors, both military and private, were merely turned loose to survive in the wild. As a result, small feral herds of Bactrian camels existed during the late 19th century in the southwest deserts of the United States. [24]

Documentaries

Military use

The Indian Army uses these camels to patrol in Ladakh. It was concluded that after carrying out trials and doing a comparative study with a single-humped camel brought from Rajasthan that the double-humped camel is better suited for the task at hand. Colonel Manoj Batra, a veterinary officer of the Indian Army, stated that the double-humped camel "are best suited for these conditions. They can carry loads of 170 kilograms (370 lb) at more than 17,000 feet (5,200 m) which is much more than the ponies that are being used as of now. They can survive without water for at least 72 hours." [25]

Footnotes

  1. ↑ A mnemonic that helps one to remember the correct English word for each is: "Bactrian" begins with "B", and "Dromedary" begins with "D"; "B" on its side has two humps, whilst "D" on its side has only one hump.

See also

Related Research Articles

<span class="mw-page-title-main">Camel</span> Genus of mammals

A camel is an even-toed ungulate in the genus Camelus that bears distinctive fatty deposits known as "humps" on its back. Camels have long been domesticated and, as livestock, they provide food and textiles. Camels are working animals especially suited to their desert habitat and are a vital means of transport for passengers and cargo. There are three surviving species of camel. The one-humped dromedary makes up 94% of the world's camel population, and the two-humped Bactrian camel makes up 6%. The wild Bactrian camel is a distinct species that is not ancestral to the domestic Bactrian camel, and is now critically endangered, with fewer than 1,000 individuals.

<span class="mw-page-title-main">Llama</span> Species of wooly domesticated mammal

The llama is a domesticated South American camelid, widely used as a meat and pack animal by Andean cultures since the pre-Columbian era.

<span class="mw-page-title-main">Feral</span> Wild-living but normally domestic animal or plant that has returned to the wild.

A feral animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species, the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in some cases, contributed to extinction of indigenous species. The removal of feral species is a major focus of island restoration.

<span class="mw-page-title-main">Camelidae</span> Family of mammals

Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, and guanacos. Camelids are even-toed ungulates classified in the order Artiodactyla, along with species including whales, pigs, deer, cattle, and antelopes.

<span class="mw-page-title-main">Dromedary</span> One-humped camel

The dromedary, also known as the dromedary camel, Arabian camel, or one-humped camel, is a large camel, of the genus Camelus, with one hump on its back.

<i>Lama</i> (genus) Genus of mammals

Lama is a genus containing the South American camelids: the wild guanaco and vicuña and the domesticated llama, alpaca, and the extinct chilihueque. Before the Spanish conquest of the Americas, llamas, alpacas, and chilihueques were the only domesticated ungulates of the continent. They were kept not only for their value as beasts of burden, but also for their flesh, hides, and wool.

<span class="mw-page-title-main">Pack animal</span> Individual or type of working animal used by humans

A pack animal, also known as a sumpter animal or beast of burden, is a working animal used to transport goods or materials by carrying them, usually on its back.

A cama is a hybrid between a male dromedary camel and a female llama, and has been produced via artificial insemination at the Camel Reproduction Centre in Dubai. The first cama was born on January 14, 1998. The aim was to create an animal capable of higher wool production than the llama, with the size and strength of a camel and a cooperative temperament.

<i>Camelops</i> Extinct genus of mammals

Camelops is an extinct genus of camel that lived in North and Central America from the middle Pliocene to the end of the Pleistocene. It is more closely related to living camels than to lamines, making it a true camel of the Camelini tribe. Its name is derived from the Ancient Greek κάμηλος and ὄψ, i.e. "camel-face". Camelops lived across western North America, ranging from the Pacific Coast to the Great Plains, southwards to Honduras and northwards to Alaska. Camelops became extinct as part of the end-Pleistocene extinction event along with most large mammals across the Americas. The extinctions followed the arrival of humans to the Americas, and evidence has been found indicating that humans butchered Camelops, suggesting that hunting may have been a factor in its extinction.

<span class="mw-page-title-main">Camel milk</span> Milk produced by female camels

Camel milk is milk from female camels. It has supported nomad and pastoral cultures since the domestication of camels millennia ago. Herders may for periods survive solely on the milk when taking the camels on long distances to graze in desert and arid environments, especially in parts of the Middle East, North Africa and the Horn of Africa. The camel dairy farming industry has grown in Australia and the United States, as an environmentally friendly alternative to cow dairy farming using a species well-adapted to arid regions.

<span class="mw-page-title-main">Alpaca</span> Domesticated species of South American camelid

The alpaca is a species of South American camelid mammal. Traditionally, alpacas are kept in herds that graze on the level heights of the Andes of Southern Peru, Western Bolivia, Ecuador, and Northern Chile. Today, alpacas may be found on farms and ranches worldwide, with thousands of animals born and raised annually. Alpacas are especially popular in North America, Europe, and Australia.

Interspecific pregnancy is the pregnancy involving an embryo or fetus belonging to another species than the carrier. Strictly, it excludes the situation where the fetus is a hybrid of the carrier and another species, thereby excluding the possibility that the carrier is the biological mother of the offspring. Strictly, interspecific pregnancy is also distinguished from endoparasitism, where parasite offspring grow inside the organism of another species, not necessarily in the womb.

<span class="mw-page-title-main">Hybrid camel</span> Hybrid between a Bactrian camel and dromedary

The hybrid camel is a domestic camelid hybrid between a Bactrian camel and dromedary. It is the offspring of a male Bactrian camel and a female dromedary.

<span class="mw-page-title-main">Camel urine</span> Liquid by-product of metabolism in camels

Camel urine is a liquid by-product of metabolism in a camel's anatomy. Urine from camels has been used in medicine for centuries, being a part of ancient Bedouin, ayurvedic and Muslim tradition. According to the World Health Organization, the use of camel urine as a medicine lacks scientific evidence. After the spread of MERS-CoV infections, the WHO urged people to refrain from drinking "raw camel milk or camel urine or eating meat that has not been properly cooked".

<span class="mw-page-title-main">Wild Bactrian camel</span> Species of camel

The wild Bactrian camel is an endangered species of camel endemic to Northwest China and southwestern Mongolia. It is closely related but not ancestral to the domestic Bactrian camel. Genetic studies have established it as a separate species which diverged from the Bactrian camel about 0.7–1.1 million years ago.

<span class="mw-page-title-main">Induced ovulation (animals)</span> Ovulation in response to an external stimulus

Induced ovulation occurs in some animal species that do not ovulate cyclically or spontaneously. Ovulation can be induced by externally-derived stimuli during or before mating, such as sperm, pheromones, or mechanical stimulation during copulation.

Camelus thomasi is an extinct species of camel from the Early-Mid Pleistocene of North Africa. It is known primarily from Tighennif (Ternifine) in Algeria. Fossils from northern Sudan and Palestine dated to the Late Pleistocene have been included under C. thomasi, but they are now considered to belong to different species, making C. thomasi a strictly Northwest African species.

Camelus knoblochi is an extinct species of camel that inhabited Eurasia during the Pleistocene epoch. One of the largest known camel species, its range spanned from Eastern Europe to Northern China.

References

  1. 1 2 Grubb, P. (2005). "Order Artiodactyla". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 645–646. ISBN   978-0-8018-8221-0. OCLC   62265494.
  2. 1 2 "Bactrian camel". EdgeofExistence.org. EDGE. 2010.
  3. ↑ "Quick Facts about Camels' Domestication History". arkbiodiv.com. 2021.
  4. 1 2 Yuan, Junxia; Hu, Jiaming; Liu, Wenhui; Chen, Shungang; Zhang, Fengli; Wang, Siren; Zhang, Zhen; Wang, Linying; Xiao, Bo; Li, Fuqiang; Hofreiter, Michael; Lai, Xulong; Westbury, Michael V.; Sheng, Guilian (May 2024). "Camelus knoblochi genome reveals the complex evolutionary history of Old World camels". Current Biology. 34 (11): 2502–2508.e5. doi:10.1016/j.cub.2024.04.050. PMID   38754423.
  5. ↑ Groves, C.; Grubb, P. (2011). Ungulate Taxonomy. Johns Hopkins University Press. p. 32. ISBN   978-1-4214-0093-8.
  6. ↑ de Buffon, C. (1791). Natural History, General and Particular. Vol. 6. London, UK: Alexander Strahan. p. 121.
  7. ↑ Smith, W.; Anthon, C. (1870). A Dictionary of Greek and Roman Antiquities (3rd ed.). New York, NY: Harper and Brothers Publishers. p.  204.
  8. ↑ Linnaeus, C. (1758). Systema Naturæ per Regna Tria Naturae. Vol. 1 (10th  ed.). Stockholm, Sweden: Laurentius salvius. p. 65.
  9. ↑ Mukasa-Mugerwa, E. (1981). The camel (Camelus dromedarius): A bibliographical review (PDF). Addis Ababa, Ethiopia: International Livestock Centre for Africa. pp. 1–147.
  10. ↑ Stanley, H.F.; Kadwell, M.; Wheeler, J.C. (1994). "Molecular evolution of the family Camelidae: A mitochondrial DNA study". Proceedings of the Royal Society B: Biological Sciences. 256 (1345): 1–6. Bibcode:1994RSPSB.256....1S. doi:10.1098/rspb.1994.0041. PMID   8008753. S2CID   40857282.
  11. ↑ Makhdoomi, Dil M.; Gazi, Mohsin A.; ul Nabi, Showkat; Ahmed, Shakeel (2013). "Morphometric studies on adult double humped camel of Ladakh, India". Emirates Journal of Food and Agriculture . Animal Science. 25 (7): 544–548. doi: 10.9755/ejfa.v25i7.15999 .
  12. ↑ "Camel fact sheet". Nature. PBS. 2020.
  13. ↑ "Bactrian camel". Dimensions.com.
  14. ↑ Chen, B.X.; Yuen, Z.X.; Pan, G.W. (1 July 1985). "Semen-induced ovulation in the bactrian camel (Camelus bactrianus)". Reproduction . 74 (2): 335–339. doi: 10.1530/jrf.0.0740335 . ISSN   0022-4251. PMID   3900379. S2CID   2219773.
  15. ↑ "Bactrian Camel". UltimateUngulate.com.
  16. ↑ "Bactrian Camel". Highland Wildlife Park. Retrieved 15 July 2022.
  17. ↑ Wand, C.; Richardson, C. (November 2009). "Replacing water with clean snow for ewes and beef cows" (PDF). OMAFRA.gov.on.ca. Ontario Ministry of Agriculture, Food, and Rural Affairs. 9–065 AGDEX 424/433. Archived from the original (PDF) on 2 April 2015. Retrieved 9 February 2017.
  18. ↑ Chen, B.X.; Yuen, Z.X. & Pan, G.W. (1985). "Semen-induced ovulation in the bactrian camel (Camelus bactrianus)" (PDF). J. Reprod. Fertil. 74 (2): 335–339. doi: 10.1530/jrf.0.0740335 . PMID   3900379.
  19. ↑ This review... Wang, Guo-Dong; Xie, Hai-Bing; Peng, Min-Sheng; Irwin, David; Zhang, Ya-Ping (1 February 2014). "Domestication Genomics: Evidence from Animals". Annual Review of Animal Biosciences . 2 (1). Annual Reviews: 65–84. doi:10.1146/annurev-animal-022513-114129. ISSN   2165-8102. PMID   25384135. ...cites this study: Wang, Zhen; Ding, Guohui; Chen, Gangliang; Sun, Yamin; Sun, Zhihong; Zhang, Heping; Wang, Lei; Hasi, S.; Zhang, Yan; Li, Jianmei; Shi, Yixiang; Xu, Ze; He, Chuan; Yu, Siriguleng; Li, Shengdi; Zhang, Wenbin; Batmunkh, Mijiddorj; Ts, Batsukh; Bat-Ireedui, Shirzana; Gao, Hongwei; Baysgalan, B.; Li, Qing; Jia, Zhi-lei; Wang, Zhaoxia; Wang, Juan; Pan, Lei; Chen, Yongcan; Ganerdene, Yaichil; Liu, Tuya; Cao, M.; He, Fei; Zha-ti, A.; Zheng, Guangyong; Qiu, Feng; Sun, Zikui; Zhao, Lele; Zhao, Wenjing; Liu, Baohong; Li, Chao; Chen, Yun-Qin; Tang, Xiaoyan; Guo, C.; Liu, Wei; Ming, Liang; Cui, Aiying; Li, Yi; Gao, Junhui; Li, Jing; Niu, Shen; Sun, Tao; Zhai, Zhengxiao; Zhang, Min; Chen, Chen; Baldan, T.; Bayaer, Tuman; Li, Yixue; Meng, H.; Bactrian Camels Genome Sequencing and Analysis Consortium (2012). "Genome sequences of wild and domestic bactrian camels". Nature Communications . 3 (1). Nature Portfolio: 1202. Bibcode:2012NatCo...3.1202T. doi:10.1038/ncomms2192. ISSN   2041-1723. PMC   3514880 . PMID   23149746. S2CID   13218949.
  20. 1 2 Burger, Pamela Anna (2016). "The history of Old World camelids in the light of molecular genetics". Tropical Animal Health and Production . 48 (5). Springer Science+Business Media: 905–913. doi:10.1007/s11250-016-1032-7. ISSN   0049-4747. PMC   4884201 . PMID   27048619.
  21. ↑ MacHugh, David E.; Larson, Greger; Orlando, Ludovic (8 February 2017). "Taming the Past: Ancient DNA and the Study of Animal Domestication". Annual Review of Animal Biosciences . 5 (1). Annual Reviews: 329–351. doi:10.1146/annurev-animal-022516-022747. ISSN   2165-8102. PMID   27813680.
  22. 1 2 Rahimdad Khan Molai Shedai; Janat ul Sindh 3rd edition 1993; Sindhi Adbi Board Jamshoro, page 20
  23. ↑ Andrea, A.J.; Levi, S.C. (2009). "Afro-Eurasian connectivity across the ages" (PDF). World System History. The Silk Road. Vol. I.
  24. ↑ Zentner, Joe. "The desert camel experiment". DesertUSA.com and Digital West Media. Retrieved 9 February 2017.
  25. ↑ Bhalla, Abhishek (19 September 2020). "Indian Army to use double-humped camels for transportation, patrolling in Ladakh". India Today.
This page is based on this Wikipedia article
Text is available under the CC BY-SA 4.0 license; additional terms may apply.
Images, videos and audio are available under their respective licenses.