Capra (genus)

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Capra
Temporal range: 2.58–0  Ma
Koze iznad St.Leonarda, Juzhni Tirol, Italija.jpg
Capra, St. Leonhard in Passeier, Italy
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Family: Bovidae
Subfamily: Caprinae
Tribe: Caprini
Genus: Capra
Linnaeus, 1758
Type species
Capra hircus
Species

See text.

Capra range map2.jpg
Approximate range of the Capra species

Capra is a genus of mammals, the goats, composed of up to nine species, including the markhor and many species known as ibexes. The domestic goat (Capra hircus) is a domesticated species derived from the wild goat (Capra aegagrus). Evidence of goat domestication dates back more than 8,500 years.

Contents

Wild goats are animals of mountain habitats. They are very agile and hardy, able to climb on bare rock and survive on sparse vegetation. They can be distinguished from the genus Ovis , which includes sheep, by the presence of scent glands close to the feet, in the groin, and in front of the eyes, and the absence of other facial glands, and by the presence of a beard in some specimens, and of hairless calluses on the knees of the forelegs. [1]

The Rocky Mountain goat is in a separate genus, Oreamnos . Present-day genetic and phenotypic differences between the Capra species are largely related to (1) discontinuity of and impeded migration between Capra populations during glacial periods, and (2) insufficient time in the postglacial period for now-adjoining Capra populations to overcome behavioral mechanisms impeding hybridization in the wild so as to erase these differences. [2]

Taxonomy

Male Nubian ibex Capra nubiana, Tierpark Hellabrunn - 03.jpg
Male Nubian ibex
Caprine heart. Caprine heart.jpg
Caprine heart.

All members of the genus Capra are bovids (members of the family Bovidae), and more specifically caprines (subfamily Caprinae). As such they are ruminants, meaning they chew the cud, and have four-chambered stomachs which play a vital role in digesting, regurgitating, and redigesting their food.

The genus has sometimes been taken to include Ovis (sheep) and Ammotragus (Barbary sheep), [3] but these are usually regarded as distinct genera, leaving Capra for ibexes. In this smaller genus, some authors have recognized only two species, the markhor on one side and all other forms included in one species on the other side. [4] Today, nine species are usually accepted: [5]

The goats of the genus Capra have complex systematic relationships, which are still not completely resolved. Recent studies based on mitochondrial DNA suggest that the Siberian ibex and the Nubian ibex represent distinct species, which are not very closely related to the physically similar Alpine ibex. The Alpine ibex forms a group with the Spanish ibex. The West Caucasian tur appears to be more closely related to the wild goat than to the East Caucasian tur. The markhor is relatively little separated from other forms—previously it had been considered to be a separate branch of the genus. [6]

Almost all wild goat species are allopatric (geographically separated)—the only geographical overlaps are the wild goat (Capra hircus) with the East Caucasian tur (Capra caucasica cylindricornis), and the markhor (Capra falconeri) with the Siberian ibex (Capra siberica). In both cases, the overlapping species do not usually interbreed in the wild, but in captivity, all Capra species can interbreed, producing fertile offspring. [7]

Species and subspecies

ImageScientific nameCommon NameDistribution
Capra caucasica1.JPG Capra caucasica West Caucasian tur Caucasus Mountains range.
Daghestanischer Tur Ostkaukasischer Steinbock Capra cylindricornis Zoo Augsburg-10 (cropped).jpg Capra cylindricornis East Caucasian tur Greater Caucasus Mountains
Markhor Schraubenziege Capra falconeri Zoo Augsburg-02.jpg Capra falconeri Markhor Central Asia, Karakoram and the Himalayas
Bezoarziege.jpg Capra aegagrus Wild goat Turkey and the Caucasus in the west to Turkmenistan, Afghanistan and Pakistan
Gorges du Verdon Goat-Rove-black 0253.jpg Capra hircus Domesticated goat Domesticated
Steinbock-P1150170.jpg Capra sibirica Siberian ibex central and northern Asia, Afghanistan, western and northern China (Primarily Xinjiang), north-western India, south-eastern Kazakhstan, Kyrgyzstan, Tajikistan, eastern Uzbekistan, Mongolia, northern Pakistan, and south-central Russia.
Cabra montes 2.jpg Capra pyrenaica Spanish ibex Iberian Peninsula
Walia ibex 2.jpg Capra walie Walia ibex Ethiopia
SteinbockGabinten.jpg Capra ibex Alpine ibex France, Italy, Switzerland, Liechtenstein, Bavaria, Austria and Slovenia
Nubian Ibex in Negev.JPG Capra nubiana Nubian ibex Algeria, Egypt, Ethiopia, Eritrea, Israel, Jordan, Lebanon, Oman, Saudi Arabia, Sudan, and Yemen

Domestication and uses

Goats used for natural weed control Goats as weed control.jpg
Goats used for natural weed control
Ibex securely climbing rocky slope Ibex in the French Vanoise National Park.JPG
Ibex securely climbing rocky slope

Along with sheep, goats were among the first domesticated animals. The domestication process started at least 10,000 years ago in what is now northern Iran. [8] Easy human access to goat hair, meat, and milk were the primary motivations. Goat skins were popularly used until the Middle Ages for water and wine bottles when traveling and camping, and in certain regions as parchment for writing.


Related Research Articles

<span class="mw-page-title-main">Caprinae</span> Subfamily of mammals

The subfamily Caprinae, also sometimes referred to as the tribe Caprini, is part of the ruminant family Bovidae, and consists of mostly medium-sized bovids. A member of this subfamily is called a caprine, or, more informally, a goat-antelope.

<span class="mw-page-title-main">Ibex</span> Type of mammal

An ibex is any of several species of wild goat , distinguished by the male's large recurved horns, which are transversely ridged in front. Ibex are found in Eurasia, North Africa and East Africa. The name ibex comes from Latin, borrowed from Iberian or Aquitanian, akin to Old Spanish bezerro "bull", modern Spanish becerro "yearling". Ranging in height from 70 to 110 centimetres (27–43 in) and weighing 90 to 120 kilograms (200–270 lb), ibex can live up to 20 years. Two closely related varieties of goats found in the wild are not usually called ibex: the markhor and the feral goat.

<span class="mw-page-title-main">Alpine ibex</span> Species of mammal

The Alpine ibex, also known as the steinbock, bouquetin, or simply ibex, is a species of wild goat that lives in the mountains of the European Alps. It is a sexually dimorphic species: males are larger and carry longer, curved horns than females. Its coat colour is typically brownish grey. Alpine ibex tend to live in steep, rough terrain near the snow line. They are also social, although adult males and females segregate for most of the year, coming together only to mate. Four distinct groups exist; adult male groups, female-offspring groups, groups of young individuals, and mixed-sex groups.

<span class="mw-page-title-main">Markhor</span> Species of mammal

The markhor is a large Capra species native to Central Asia, the Karakoram, and the Himalayas. It is listed on the IUCN Red List as Near Threatened since 2015.

<span class="mw-page-title-main">Kri-kri</span> Feral goat

The kri-kri, sometimes called the Cretan goat, Agrimi, or Cretan Ibex, is a feral goat inhabiting the Eastern Mediterranean, previously considered a subspecies of wild goat. The kri-kri today is found only in Greece on three small islands just off the shore of Crete. The kri-kri can also be found on the island of Sapientza which was brought to the island in great numbers, in order to protect the species from extinction.

<span class="mw-page-title-main">Wild goat</span> Species of mammal

The wild goat is a wild goat species, inhabiting forests, shrublands and rocky areas ranging from Turkey and the Caucasus in the west to Turkmenistan, Afghanistan and Pakistan in the east. It has been listed as near threatened on the IUCN Red List and is threatened by destruction and degradation of habitat.

<span class="mw-page-title-main">West Caucasian tur</span> Species of mammal

The West Caucasian tur is a mountain-dwelling goat-antelope native to the western half of the Caucasus Mountains range, in Georgia and European Russia. It is listed as Endangered on the IUCN Red List, as the wild population is estimated to be between 5,000 and 6,000 individuals.

<span class="mw-page-title-main">Nubian ibex</span> Species of mammal

The Nubian ibex is a desert-dwelling goat species found in mountainous areas of northern and northeast Africa, and the Middle East. It was historically considered to be a subspecies of the Alpine ibex, but is now considered a distinct species. The wild population is estimated at 4,500 mature individuals.

The Naltar Wildlife Sanctuary is a protected area located in the Naltar Valley near Nomal, in Gilgit-Baltistan, Pakistan. The Sanctuary was created on 22 November 1975 and consists of a steep-sided forested valley with high mountains on either side. A small number of Astor markhor are found here as well as other large mammals.

<span class="mw-page-title-main">Caucasus mixed forests</span> Ecoregion

The Caucasus mixed forests is a temperate broadleaf and mixed forests ecoregion in the Caucasus Mountains, as well as the adjacent Lesser Caucasus range and the eastern end of the Pontic Mountains.

Afghanistan has long been known for diverse wildlife. Many of the larger mammals in the country are categorized by the International Union for Conservation of Nature (IUCN) as globally threatened. These include the snow leopard, Marco Polo sheep, Siberian musk deer, markhor, urial, and the Asiatic black bear. Other species of interest are the ibex, the gray wolf, and the brown bear, striped hyenas, and numerous bird of prey species. Most of the Marco Polo sheep and ibex are being poached for food, whereas wolves, snow leopards and bears are being killed for damage prevention.

<span class="mw-page-title-main">Shahdag National Park</span> Historic site in Oğuz Rayon , Şamaxı Rayon

Shahdag National Park — is a national park of Azerbaijan. It was established on an area of 115,900 hectares (1,159 km2) in the Quba Rayon, Qusar Rayon, İsmayıllı Rayon, Qəbələ Rayon, Oğuz Rayon and Şamaxı Rayon administrative districts on December 8, 2006. It was enlarged by presidential decree on July 8, 2010 from 115,900 hectares (1,159 km2) to 130,508.1 hectares (1,305.081 km2).

<span class="mw-page-title-main">Bezoar ibex</span> Subspecies of goat

The bezoar ibex is a wild goat subspecies that is native to the montane forested areas in the Caucasus and the Zagros Mountains.

Sarcocystis is a genus of parasitic Apicomplexan alveolates. Species in this genus infect reptiles, birds and mammals. The name is derived from Greek: sarkos = flesh and kystis = bladder.

<span class="mw-page-title-main">Bovidae in Chinese mythology</span>

Bovidae in Chinese mythology include various myths and legends about a group of biologically distinct animals which form important motifs within Chinese mythology. There are many myths about the animals modernly classified as Bovidae, referring to oxen, sheep, goats, and mythological types such as "unicorns". Chinese mythology refers to those myths found in the historical geographic area of China, a geographic area which has evolved or changed somewhat through history. Thus this includes myths in Chinese and other languages, as transmitted by Han Chinese as well as other ethnic groups. There are various motifs of animals of the Bovidae biological family in Chinese mythology. These have often served as allusions in poetry and other literature. Some species are also used in the traditional Chinese calendar and time-keeping system.

<span class="mw-page-title-main">Pshav-Khevsureti National Park</span> National park in Georgia

Pshav-Khevsureti National Park is in Dusheti Municipality, Mtskheta-Mtianeti region of Georgia. Park has been established by direct involvement of World Wide Fund for Nature.

<span class="mw-page-title-main">Domestication of the goat</span>

Goat evolution is the process by which domestic goats came to exist through evolution by natural selection. Wild goats — medium-sized mammals which are found in noticeably harsh environments, particularly forests and mountains, in the Middle East and Central Asia — were one of the first species domesticated by modern humans, with the date of domestication generally considered to be 8,000 BCE. Goats are part of the family Bovidae, a broad and populous group which includes a variety of ruminants such as bison, cows and sheep. Bovids all share many traits, such as hooves and a herbivorous diet and all males, along with many females, have horns. Bovids began to diverge from deer and giraffids during the early Miocene epoch. The subfamily Caprinae, which includes goats, ibex and sheep, are considered to have diverged from the rest of Bovidae as early as the late Miocene, with the group reaching its greatest diversity in the ice ages.

References

  1. Parrini, F.; et al. (2009). "Capra ibex (Artiodactyla: Bovidae)". Mammalian Species. 830: 1–12. doi: 10.1644/830.1 .
  2. Gavashelishvili, A.; Yarovenko, Y. A.; Babayev, E. A.; Mikeladze, G.; Gurielidze, Z.; Dekanoidze, D.; Kerdikoshvili, N.; Ninua, L.; Paposhvili, N. (2018). "Modeling the distribution and abundance of eastern tur (Capra cylindricornis) in the Caucasus". Journal of Mammalogy. 99 (4): 885–897. doi: 10.1093/jmammal/gyy056 .
  3. Ansell, W. F. H. 1972. Order Artiodactyla. Part 15. Pp. 1–84, in The mammals of Africa: An identification manual (J. Meester and H. W. Setzer, eds.) [issued 2 May 1972]. Smithsonian Institution Press, Washington, D.C., not continuously paginated. (quoted in Grubb, P. (2005). Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore, MD: Johns Hopkins University Press. ISBN   0-8018-8221-4. OCLC   62265494.)
  4. Haltenorth, T. 1963. Klassifikation der Säugetiere: Artiodactyla I. Handbuch der Zoologie, 8(32):1–167 (quoted in Grubb, P. (2005). Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore, MD: Johns Hopkins University Press. ISBN   0-8018-8221-4. OCLC   62265494.)
  5. Nathalie Pidancier, Steve Jordan, Gordon Luikart, Pierre Taberlet: Evolutionary history of the genus Capra (Mammalia, Artiodactyla): Discordance between mitochondrial DNA and Y-chromosome. Molecular Phylogenetics and Evolution 40 (2006) 739–749 online
  6. Phylogenetic Reconstructions in the Genus Capra (Bovidae, Artiodactyla) Based on the Mitochondrial DNA Analysis. Russian Journal of Genetics, 2007, Vol. 43, No. 2, pp. 181–189. online
  7. V. G. Heptner: Mammals of the Sowjetunion Vol. I UNGULATES. Leiden, New York, 1989 ISBN   90-04-08874-1
  8. Melinda A. Zeder, Brian Hesse: The Initial Domestication of Goats (Capra hircus) in the Zagros Mountains 10,000 Years Ago. Science 24 March 2000: Vol. 287. no. 5461, pp. 2254–2257 online abstract