|Domestic llama and alpaca range|
Camelus glamaLinnaeus, 1758
The llama ( // ; Spanish pronunciation: [ˈʎama] ) (Lama glama) is a domesticated South American camelid, widely used as a meat and pack animal by Andean cultures since the Pre-Columbian era.
Llamas are very social animals and live with others as a herd. Their wool is very soft and lanolin-free. Llamas can learn simple tasks after a few repetitions. When using a pack, they can carry about 25 to 30% of their body weight for 8 to 13 km (5–8 miles).The name llama (in the past also spelled "lama" or "glama") was adopted by European settlers from native Peruvians.
Llamas appear to have originated from the central plains of North America about 40 million years ago. They migrated to South America about three million years ago during the Great American Interchange. By the end of the last ice age (10,000–12,000 years ago), camelids were extinct in North America.As of 2007, there were over seven million llamas and alpacas in South America, and due to importation from South America in the late 20th century, there are now over 158,000 llamas and 100,000 alpacas in the United States and Canada.
Lamoids, or llamas (as they are more generally known as a group), consist of the vicuña (Vicugna vicugna, prev. Lama vicugna), guanaco (Lama guanicoe), Suri alpaca, and Huacaya alpaca (Vicugna pacos, prev. Lama guanicoe pacos), and the domestic llama (Lama glama). Guanacos and vicuñas live in the wild, while llamas and alpacas exist only as domesticated animals.Although early writers compared llamas to sheep, their similarity to the camel was soon recognized. They were included in the genus Camelus along with alpaca in the Systema Naturae (1758) of Carl Linnaeus. They were, however, separated by Georges Cuvier in 1800 under the name of lama along with the guanaco. DNA analysis has confirmed that the guanaco is the wild ancestor of the llama, while the vicuña is the wild ancestor of the alpaca; the latter two were placed in the genus Vicugna.
The genera Lama and Vicugna are, with the two species of true camels, the sole existing representatives of a very distinct section of the Artiodactyla or even-toed ungulates, called Tylopoda, or "bump-footed", from the peculiar bumps on the soles of their feet. The Tylopoda consist of a single family, the Camelidae, and shares the order Artiodactyla with the Suina (pigs), the Tragulina (chevrotains), the Pecora (ruminants), and the Whippomorpha (hippos and cetaceans, which belong to Artiodactyla from a cladistic, if not traditional, standpoint). The Tylopoda have more or less affinity to each of the sister taxa, standing in some respects in a middle position between them, sharing some characteristics from each, but in others showing special modifications not found in any of the other taxa.[ citation needed ]
The 19th-century discoveries of a vast and previously unexpected extinct Paleogene fauna of North America, as interpreted by paleontologists Joseph Leidy, Edward Drinker Cope, and Othniel Charles Marsh, aided understanding of the early history of this family.[ citation needed ] Llamas were not always confined to South America; abundant llama-like remains were found in Pleistocene deposits in the Rocky Mountains and in Central America. Some of the fossil llamas were much larger than current forms. Some species remained in North America during the last ice ages. North American llamas are categorized as a single extinct genus, Hemiauchenia . Llama-like animals would have been a common sight 25,000 years ago, in modern-day California, Texas, New Mexico, Utah, Missouri, and Florida.
The camelid lineage has a good fossil record. Camel-like animals have been traced from the thoroughly differentiated, modern species back through early Miocene forms. Their characteristics became more general, and they lost those that distinguished them as camelids; hence, they were classified as ancestral artiodactyls.[ citation needed ] No fossils of these earlier forms have been found in the Old World, indicating that North America was the original home of camelids, and that Old World camels crossed over via the Bering Land Bridge. The formation of the Isthmus of Panama three million years ago allowed camelids to spread to South America as part of the Great American Interchange, where they evolved further. Meanwhile, North American camelids died out at the end of the Pleistocene.
A full-grown llama can reach a height of 1.7 to 1.8 m (5 ft 7 in to 5 ft 11 in) at the top of the head, and can weigh between 130 and 200 kg (290 and 440 lb). At birth, a baby llama (called a cria ) can weigh between 9 and 14 kg (20 and 31 lb). Llamas typically live for 15 to 25 years, with some individuals surviving 30 years or more. [ better source needed ]
The following characteristics apply especially to llamas. Dentition of adults:-incisors 1/3 canines 1/1, premolars 2/2, molars 3/2; total 32. In the upper jaw, a compressed, sharp, pointed laniariform incisor near the hinder edge of the premaxilla is followed in the male at least by a moderate-sized, pointed, curved true canine in the anterior part of the maxilla.The isolated canine-like premolar that follows in the camels is not present. The teeth of the molar series, which are in contact with each other, consist of two very small premolars (the first almost rudimentary) and three broad molars, constructed generally like those of Camelus. In the lower jaw, the three incisors are long, spatulate, and procumbent; the outer ones are the smallest. Next to these is a curved, suberect canine, followed after an interval by an isolated minute and often deciduous simple conical premolar; then a contiguous series of one premolar and three molars, which differ from those of Camelus in having a small accessory column at the anterior outer edge.
The skull generally resembles that of Camelus, the larger brain-cavity and orbits and less-developed cranial ridges being due to its smaller size. The nasal bones are shorter and broader, and are joined by the premaxilla.
The ears are rather long and slightly curved inward, characteristically known as "banana" shaped. There is no dorsal hump. The feet are narrow, the toes being more separated than in the camels, each having a distinct plantar pad. The tail is short, and fibre is long, woolly and soft.
In essential structural characteristics, as well as in general appearance and habits, all the animals of this genus very closely resemble each other, so whether they should be considered as belonging to one, two, or more species is a matter of controversy among naturalists.
The question is complicated by the circumstance of the great majority of individuals that have come under observation being either in a completely or partially domesticated state. Many are also descended from ancestors that have previously been domesticated, a state that tends to produce a certain amount of variation from the original type. The four forms commonly distinguished by the inhabitants of South America are recognized as distinct species, though with difficulties in defining their distinctive characteristics.
The llama and alpaca are only known in the domestic state, and are variable in size and of many colors, being often white, brown, or piebald. Some are grey or black. The guanaco and vicuña are wild, the former being endangered, and of a nearly uniform light-brown color, passing into white below. They certainly differ from each other, the vicuña being smaller, more slender in its proportions, and having a shorter head than the guanaco. The vicuña lives in herds on the bleak and elevated parts of the mountain range bordering the region of perpetual snow, amidst rocks and precipices, occurring in various suitable localities throughout Peru, in the southern part of Ecuador, and as far south as the middle of Bolivia. Its manners very much resemble those of the chamois of the European Alps; it is as vigilant, wild, and timid. The fiber is extremely delicate and soft, and highly valued for the purposes of weaving, but the quantity that each animal produces is minimal. Alpacas are descended from wild vicuna ancestors, while domesticated llamas are descended from wild guanaco ancestors, though a considerable amount of hybridization between the two species has occurred.
Differential characteristics between llamas and alpacas include the llama's larger size, longer head, and curved ears. Alpaca fiber is generally more expensive, but not always more valuable. Alpacas tend to have a more consistent color throughout the body. The most apparent visual difference between llamas and camels is that camels have a hump or humps and llamas do not.
Llamas are not ruminants, pseudo-ruminants, or modified ruminants.They have a complex stomach with several compartments that allows them to consume lower quality, high cellulose foods. The stomach compartments allow for fermentation of tough food stuffs, followed by regurgitation and re-chewing. Ruminants have four compartments (cows, sheep, goats), whereas llamas have only three stomach compartments: the rumen, omasum, and abomasum.
In addition, the llama (and other camelids) have an extremely long and complex large intestine (colon). The large intestine's role in digestion is to reabsorb water, vitamins and electrolytes from food waste that is passing through it. The length of the llama's colon allows it to survive on much less water than other animals. This is a major advantage in arid climates where they live.
Llamas have an unusual reproductive cycle for a large animal. Female llamas are induced ovulators.Through the act of mating, the female releases an egg and is often fertilized on the first attempt. Female llamas do not go into estrus ("heat").
Like humans, llama males and females mature sexually at different rates. Females reach puberty at about 12 months old; males do not become sexually mature until around three years of age.
Llamas mate with in a kush (lying down) position, which is fairly unusual in a large animal. They mate for an extended time (20–45 minutes), also unusual in a large animal.
The gestation period of a llama is 11.5 months (350 days). Dams (female llamas) do not lick off their babies, as they have an attached tongue that does not reach outside of the mouth more than half an inch (1.3 cm). Rather, they will nuzzle and hum to their newborns.
A cria (from Spanish for "baby") is the name for a baby llama, alpaca, vicuña, or guanaco. Crias are typically born with all the females of the herd gathering around, in an attempt to protect against the male llamas and potential predators. Llamas give birth standing. Birth is usually quick and problem-free, over in less than 30 minutes. Most births take place between 8 am and noon, during the warmer daylight hours. This may increase cria survival by reducing fatalities due to hypothermia during cold Andean nights. This birthing pattern is speculated to be a continuation of the birthing patterns observed in the wild. Crias are up and standing, walking and attempting to suckle within the first hour after birth. Crias are partially fed with llama milk that is lower in fat and salt and higher in phosphorus and calcium than cow or goat milk. A female llama will only produce about 60 ml (2.1 imp fl oz) of milk at a time when she gives milk, so the cria must suckle frequently to receive the nutrients it requires.
In harem mating, the male is left with females most of the year.
For field mating, a female is turned out into a field with a male llama and left there for some period of time. This is the easiest method in terms of labor, but the least useful in terms of prediction of a likely birth date. An ultrasound test can be performed, and together with the exposure dates, a better idea of when the cria is expected can be determined.
Hand mating is the most efficient method, but requires the most work on the part of the human involved. A male and female llama are put into the same pen and mating is monitored. They are then separated and re-mated every other day until one or the other refuses the mating. Usually, one can get in two matings using this method, though some stud males routinely refuse to mate a female more than once. The separation presumably helps to keep the sperm count high for each mating and also helps to keep the condition of the female llama's reproductive tract more sound. If the mating is not successful within two to three weeks, the female is mated again.
This section contains instructions, advice, or how-to content . (December 2016)
Llamas should be tested for pregnancy after mating at two to three, six and at least 12 weeks.
Spit testing with an intact male is generally free and is usually accurate. However, some hormonal conditions in females can make them reject a male when they are in fact not pregnant, and, more rarely, accept a male when they are pregnant. Progesterone tests can give a high reading in some females with a hormonal problem that are in fact not pregnant. Neither of the previous methods, nor palpation, can give a reasonably accurate idea of the age of the fetus, while an ultrasound procedure can. In addition, an ultrasound procedure can distinguish between pregnancy and misleading physical conditions, or between a live and dead fetus. The disadvantages of an ultrasound procedure are cost, some training in the use of ultrasound equipment is required, and not all veterinarians have the equipment needed to perform the examination.
Options for feeding llamas are quite wide; a wide variety of commercial and farm-based feeds are available. The major determining factors include feed cost, availability, nutrient balance and energy density required. Young, actively growing llamas require a greater concentration of nutrients than mature animals because of their smaller digestive tract capacities.
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Llamas that are well-socialized and trained to halter and lead after weaning are very friendly and pleasant to be around. They are extremely curious and most will approach people easily. However, llamas that are bottle-fed or over-socialized and over-handled as youth will become extremely difficult to handle when mature, when they will begin to treat humans as they treat each other, which is characterized by bouts of spitting, kicking and neck wrestling.
Llamas have started showing up in nursing homes and hospitals as certified therapy animals. Rojo the Llama, located in the Pacific Northwest was certified in 2008. The Mayo Clinic says animal-assisted therapy can reduce pain, depression, anxiety, and fatigue. This type of therapy is growing in popularity, and there are several organizations throughout the United States that participate.
When correctly reared, llamas spitting at a human is a rare thing. Llamas are very social herd animals, however, and do sometimes spit at each other as a way of disciplining lower-ranked llamas in the herd. A llama's social rank in a herd is never static. They can always move up or down in the social ladder by picking small fights. This is usually done between males to see which will become dominant. Their fights are visually dramatic, with spitting, ramming each other with their chests, neck wrestling and kicking, mainly to knock the other off balance. The females are usually only seen spitting as a means of controlling other herd members. One may determine how agitated the llama is by the materials in the spit. The more irritated the llama is, the further back into each of the three stomach compartments it will try to draw materials from for its spit.
While the social structure might always be changing, they live as a family and they do take care of each other. If one notices a strange noise or feels threatened, an alarm call - a loud, shrill sound which rhythmically rises and falls - is sent out and all others become alert. They will often hum to each other as a form of communication.
The sound of the llama making groaning noises or going "mwa" (/mwaʰ/) is often a sign of fear or anger. Unhappy or agitated llamas will lay their ears back, while ears being perked upwards is a sign of happiness or curiosity.
An "orgle" is the mating sound of a llama or alpaca, made by the sexually aroused male. The sound is reminiscent of gargling, but with a more forceful, buzzing edge. Males begin the sound when they become aroused and continue throughout the act of procreation – from 15 minutes to more than an hour.
Using llamas as livestock guards in North America began in the early 1980s, and some sheep producers have used llamas successfully since then. Some would even use them to guard their smaller cousins, the alpaca.They are used most commonly in the western regions of the United States, where larger predators, such as coyotes and feral dogs, are prevalent. Typically, a single gelding (castrated male) is used.
Research suggests the use of multiple guard llamas is not as effective as one. Multiple males tend to bond with one another, rather than with the livestock, and may ignore the flock. A gelded male of two years of age bonds closely with its new charges and is instinctively very effective in preventing predation. Some llamas appear to bond more quickly to sheep or goats if they are introduced just prior to lambing. Many sheep and goat producers indicate a special bond quickly develops between lambs and their guard llama and the llama is particularly protective of the lambs.
Using llamas as guards has reduced the losses to predators for many producers. The value of the livestock saved each year more than exceeds the purchase cost and annual maintenance of a llama. Although not every llama is suited to the job, most are a viable, nonlethal alternative for reducing predation, requiring no training and little care. [ better source needed ]
Scholar Alex Chepstow-Lusty has argued that the switch from a hunter-gatherer lifestyle to widespread agriculture was only possible because of the use of llama dung as fertilizer.
The Moche people frequently placed llamas and llama parts in the burials of important people, as offerings or provisions for the afterlife.The Moche culture of pre-Columbian Peru depicted llamas quite realistically in their ceramics.
In the Inca Empire, llamas were the only beasts of burden, and many of the people dominated by the Inca had long traditions of llama herding. For the Inca nobility, the llama was of symbolic significance, and llama figures were often buried with the dead.In South America, llamas are still used as beasts of burden, as well as for the production of fiber and meat.
The Inca deity Urcuchillay was depicted in the form of a multicolored llama.
Carl Troll has argued that the large numbers of llamas found in the southern Peruvian highlands were an important factor in the rise of the Inca Empire.It is worth considering the maximum extent of the Inca Empire roughly coincided with the greatest distribution of alpacas and llamas in Pre-Hispanic America. The link between the Andean biomes of puna and páramo, llama pastoralism and the Inca state is a matter of research.
One of the main uses for llamas at the time of the Spanish conquest was to bring down ore from the mines in the mountains.Gregory de Bolivar estimated that in his day, as many as 300,000 were employed in the transport of produce from the Potosí mines alone, but since the introduction of horses, mules, and donkeys, the importance of the llama as a beast of burden has greatly diminished.
According to Juan Ignacio Molina, the Dutch captain Joris van Spilbergen observed the use of hueques (possibly a llama type) by native Mapuches of Mocha Island as plow animals in 1614.
In Chile hueque populations declined towards extinction in the 16th and 17th century being replaced by European livestock.The causes of its extinction are not clear but it is known that the introduction of sheep caused some competition among both domestic species. Anecdotal evidence of the mid-17th century show that both species coexisted but suggests that there were many more sheep than hueques. The decline of hueques reached a point in the late 18th century when only the Mapuche from Mariquina and Huequén next to Angol raised the animal.
Llamas were first imported into the US in the late 1800s as zoo exhibits. Restrictions on importation of livestock from South America due to hoof and mouth disease, combined with lack of commercial interest, resulted in the number of llamas staying low until the late 20th century. In the 1970s, interest in llamas as livestock began to grow, and the number of llamas increased as farmers bred and produced an increasing number of animals.Both the price and number of llamas in the US climbed rapidly in the 1980s and 1990s. With little market for llama fiber or meat in the US, and the value of guard llamas limited, the primary value in llamas was in breeding more animals, a classic sign of a speculative bubble in agriculture. By 2002, there were almost 145,000 llamas in the US according to the US Department of Agriculture, and animals sold for as much as $220,000. However, the lack of any end market for the animals resulted in a crash in both llama prices and the number of llamas; the Great Recession further dried up investment capital, and the number of llamas in the US began to decline as fewer animals were bred and older animals died of old age. By 2017, the number of llamas in the US had dropped below 40,000. A similar speculative bubble was experienced with the closely related alpaca, which burst shortly after the llama bubble.
Llamas have a fine undercoat, which can be used for handicrafts and garments. The coarser outer guard hair is used for rugs, wall-hangings and lead ropes. The fiber comes in many different colors ranging from white or grey to reddish-brown, brown, dark brown and black.
|Angora rabbit (Angora wool)||13|
|Cashmere goat (Cashmere wool)||15–19|
|Yak (Yak fiber)||15–19|
|Camel (Camel hair)||16–25|
|Angora goat (Mohair)||25–45|
|Wikispecies has information related to Lama glama|
|Wikimedia Commons has media related to Llama .|
A camel is an even-toed ungulate in the genus Camelus that bears distinctive fatty deposits known as "humps" on its back. Camels have long been domesticated and, as livestock, they provide food and textiles. Camels are working animals especially suited to their desert habitat and are a vital means of transport for passengers and cargo. There are three surviving species of camel. The one-humped dromedary makes up 94% of the world's camel population, and the two-humped Bactrian camel makes up 6%. The Wild Bactrian camel is a separate species and is now critically endangered.
The even-toed ungulates are ungulates – hoofed animals – which bear weight equally on two of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, odd-toed ungulates bear weight on one of the five toes: the third toe. Another difference between the two is that even-toed ungulates digest plant cellulose in one or more stomach chambers rather than in their intestine as the odd-toed ungulates do.
Tylopoda is a suborder of terrestrial herbivorous even-toed ungulates belonging to the order Artiodactyla. They are found in the wild in their native ranges of South America and Asia, while Australian feral camels are introduced. The group has a long fossil history in North America and Europe. Tylopoda appeared during the Eocene around 46.2 million years ago.
Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, and guanacos. Camelids are even-toed ungulates classified in the order Cetartiodactyla, along with pigs, whales, deer, cattle, antelope, and many others.
The dromedary, also called the Arabian camel, is a large, even-toed ungulate with one hump on its back. It is the tallest of the three species of camel; adult males stand 1.8–2 m (5.9–6.6 ft) at the shoulder, while females are 1.7–1.9 m (5.6–6.2 ft) tall. Males typically weigh between 400 and 600 kg, and females weigh between 300 and 540 kg. The species' distinctive features include its long, curved neck, narrow chest, a single hump, and long hairs on the throat, shoulders and hump. The coat is generally a shade of brown. The hump, 20 cm (7.9 in) tall or more, is made of fat bound together by fibrous tissue.
The Bactrian camel is a large, even-toed ungulate native to the steppes of Central Asia. It has two humps on its back, in contrast to the single-humped dromedary camel. Its population of two million exists mainly in the domesticated form. Their name comes from the ancient historical region of Bactria.
The guanaco is a camelid native to South America, closely related to the llama. Its name comes from the Quechua word huanaco. Young guanacos are called chulengos.
The vicuña or vicuna is one of the two wild South American camelids which live in the high alpine areas of the Andes, the other being the guanaco. They were named after the eponymous town, which in turn took its name from Joaquín Vicuña, a Chilean politician of Basque origin. Vicuñas are relatives of the llama, and are now believed to be the wild ancestor of domesticated alpacas, which are raised for their coats. Vicuñas produce small amounts of extremely fine wool, which is very expensive because the animal can only be shorn every three years and has to be caught from the wild. When knitted together, the product of the vicuña's wool is very soft and warm. The Inca valued vicuñas highly for their wool, and it was against the law for anyone but royalty to wear vicuña garments; today, the vicuña is the national animal of Peru and appears in the Peruvian coat of arms.
Lama is a genus containing two South American camelids, the wild guanaco and the domesticated llama. This genus is closely allied to the wild vicuña and domesticated alpaca of the genus Vicugna. Before the Spanish conquest of the Americas, llamas and alpacas were the only domesticated ungulates of the continent. They were kept not only for their value as beasts of burden, but also for their flesh, hides, and wool.
A cama is a hybrid between a male dromedary camel and a female llama, and has been produced via artificial insemination at the Camel Reproduction Centre in Dubai. The first cama was born on January 14, 1998. The aim was to create an animal capable of higher wool production than the llama, with the size and strength of a camel and a cooperative temperament.
Vicugna is a genus containing two South American camelids, the vicuña and the alpaca.
A guard llama is a llama, guanaco, alpaca or hybrid that is used in farming to protect sheep, goats, hens or other livestock from coyotes, dogs, foxes and other predators. In the past, a single gelded (castrated) male was recommended. In more recent years it has been discovered that single, unbred females make better and safer guardians. They also do not pose the risk of attempting copulation or chasing the smaller livestock.
Inca cuisine originated in pre-Columbian times within the Inca civilization from the 13th to the 16th century. The Inca civilization stretched across many regions, and so there was a great diversity of plants and animals used for food, many of which remain unknown outside Peru. The most important staples were various tubers, roots, and grains. Maize was of high prestige, but could not be grown as extensively as it was further north. The most common sources of meat were guinea pigs and llamas, and dried fish was common.
A cria is a juvenile llama, alpaca, vicuña, or guanaco.
The alpaca is a species of South American camelid descended from the vicuña. It is similar to, and often confused with, the llama. However, alpacas are often noticeably smaller than llamas. The two animals are closely related and can successfully cross-breed. Alpacas and llamas are related to the guanaco. There are two breeds of alpaca: the Suri alpaca and the Huacaya alpaca.
Andrew Charles ("Andy") Tillman is one of the founders of the llama industry in the United States. He is an expert on llama and alpaca health, selective breeding, and marketing. Tillman is the co-founder of the International Llama Association, and he wrote the halter-class guidelines for the American Llama Show Association. His book, Speechless Brothers, was the first comprehensive study of llama husbandry published in the United States.
Lamini is a tribe of the subfamily Camelinae. It contains two extant genera with four species, all exclusively from South America: llamas, alpacas, vicuñas, and guanacos. The former two are domesticated species, while the latter two are only found in the wild. None display sexual dimorphism. The four species can interbreed and produce fertile offspring. Additionally, there are two extinct genera known from the fossil record.
Chilihueque or Hueque is a South American camelid variety or species that existed in central and south-central Chile in Pre-Hispanic and colonial times. There are two main hypotheses on their status among South American camelids: the first one suggests that they are locally domesticated guanacos and the second that they are a variety of llamas brought from the north into south-central Chile. The alpaca has also been suggested as a possible identity.
Huacaya is the one of the two breeds that make up the species Vicugna pacos, commonly known as the alpaca. The other breed is the Suri. It lives on the Altiplano plateau in the Andes at up to 4,000 m above sea level. Its natural range encompasses four South American countries.