Titanotylopus Temporal range: | |
---|---|
T. nebraskensis skeleton at Natural History Museum Los Angeles | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Artiodactyla |
Family: | Camelidae |
Tribe: | Camelini |
Genus: | † Titanotylopus Barbour & Schultz, 1934 [1] |
Species | |
†Titanotylopus nebraskensis Contents | |
Synonyms | |
|
Titanotylopus is an extinct genus of camel (tribe Camelini), endemic to North America from the late Hemphillian stage of the Miocene through the Irvingtonian stage of the Pleistocene. [2] It was one of the last surviving North American camels; after its extinction, only Camelops remained.
Its name is derived from the Greek words Τιτάν, τύλος and πούς — "Titan", "knob" and "foot"; thus, "giant knobby-foot".
Titanotylopus is distinguished from other early large camelids by its large upper canines amongst other distinguishing dental characteristics, and absence of lacrimal vacuities in the skull. Unlike the smaller, contemporaneous Camelops , Titanotylopus had relatively broad second phalanges, suggesting that it had true padded "cameltoes", like modern camels. [3]
The species Titanotylopus spatulus was characterized by broad, spatula-like incisors. It has been found at Grand View, Red Light fauna of the Love Formation, Hudspeth County, Texas, Donnelly Ranch, White Rock, Kansas, Mullen II (Kansas), Sandahl Local Fauna (Nebraska), Vallecito Creek, Colorado and 111 Ranch, Arizona in North America.
Titanotylopus possessed long and massive limbs, a comparatively small braincase, and a convex slope between the eyes. It reached a shoulder height of 3–3.5 m (9.8–11.5 ft) and a weight of 2,500 kg (5,500 lb). [4] [5]
Like modern camels, it possessed a hump for fat storage; evidence for this is provided by the long neural spines on its thoracic vertebrae. [3]
While some authors have considered Gigantocamelus and Titanotylopus to be congeneric, others have maintained them separately. Voorhies and Corner, based on previously unreported material, documented that the two are indeed worthy of separate generic status. Harrison (1985) followed Voorhies and Corner in advocating the use of Titanotylopus for only T. nebraskensis, based on a lower jaw, and Gigantocamelus for G. spatulus, which includes G. fricki. There is a clear difference between the proximal phalanx of specimens assigned to Gigantocamelus and to Titanotylopus, based on skeletons associated with skull material.[ citation needed ]
Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, and guanacos. Camelids are even-toed ungulates classified in the order Artiodactyla, along with species including whales, pigs, deer, cattle, and antelopes.
Aepycamelus is an extinct genus of camelids that lived during the Miocene 20.6–4.9 million years ago, existing for about 15.7 million years. Its name is derived from the Homeric Greek αἰπύς, "high and steep" and κάμηλος – "camel"; thus, "high camel"; alticamelus in Latin.
Camelops is an extinct genus of camels that lived in North and Central America, ranging from Alaska to Honduras, from the middle Pliocene to the end of the Pleistocene. It is more closely related to the Old World dromedary and bactrian and wild bactrian camels than to the New World guanaco, vicuña, alpaca and llama; making it a true camel of the Camelini tribe. Its name is derived from the Ancient Greek κάμηλος and ὄψ, i.e. "camel-face".
Macrauchenia is an extinct genus of large ungulate native to South America from the late Pliocene to the end of the Pleistocene. It is a member of the extinct order Litopterna, a group of South American native ungulates distinct from the two orders which contain all living ungulates which had been present in South America since the early Cenozoic, over 60 million years ago, prior to the arrival of living ungulates in South America around 2.5 million years ago as part of the Great American Interchange. The bodyform of Macrauchenia has been described as similar to a camel, being one of the largest known litopterns, with an estimated body mass of around 1 tonne. The genus gives its name to its family, Macraucheniidae, which like Macrauchenia typically had long necks and three toed feet, as well a retracted nasal region, which in Macrauchenia manifests as the nasal opening being on the top of the skull behind the eye sockets. This has historically been argued to correspond to the presence of a tapir-like proboscis, but some recent authors suggest a moose-like prehensile lip or a saiga antelope-like nose to filter dust are more likely.
Hayoceros is an extinct genus of the artiodactyl family Antilocapridae, endemic to North America during the Pleistocene epoch, existing for about 1.5 million years.
Palorchestes is an extinct genus of large terrestrial, herbivorous Australian marsupial of the family Palorchestidae, living from the Miocene through to the Late Pleistocene. Like other palorchestids, it had highly retracted nasal region suggesting that it had a prehensile lip, as well as highly unusual clawed forelimbs that were used to grasp vegetation.
Chapalmalania is an extinct genus of procyonid from the Late Miocene or Early Pliocene to Early Pleistocene of Argentina, Venezuela, and Colombia.
Merycoidodontoidea, previously known as "oreodonts" or "ruminating hogs," are an extinct superfamily of prehistoric cud-chewing artiodactyls with short faces and fang-like canine teeth. As their name implies, some of the better known forms were generally hog-like, and the group has traditionally been placed within the Suina, though some recent work suggests they may have been more closely related to camels. "Oreodont" means "mountain teeth," referring to the appearance of the molars. Most oreodonts were sheep-sized, though some genera grew to the size of cattle. They were heavy-bodied, with short four-toed hooves and comparatively long tails.
Merycoidodon is an extinct genus of herbivorous artiodactyl of the family Merycoidodontidae, more popularly known by the name Oreodon. It was endemic to North America during the Middle Eocene to Middle Miocene existing for approximately 30 million years.
Metridiochoerus is an extinct genus of swine known from the Pliocene and Pleistocene of Africa. It is also known as the giant warthog.
Synthetoceras tricornatus is a large, extinct protoceratid, endemic to North America during the Late Miocene, 10.3—5.3 Ma, existing for approximately 5 million years. Fossils have been recovered from Nebraska and Texas.
Poebrotherium is an extinct genus of camelid, endemic to North America. They lived from the Eocene to Miocene epochs, 46.3—13.6 mya, existing for approximately 32 million years.
Sivatherium is an extinct genus of giraffids that ranged throughout Africa to the Indian subcontinent. The species Sivatherium giganteum is, by weight, one of the largest giraffids known, and also one of the largest ruminants of all time.
Paramylodon is an extinct genus of ground sloth of the family Mylodontidae endemic to North America during the Pliocene through Pleistocene epochs, living from around ~4.9 Mya–12,000 years ago.
Mount Blanco is a small white hill — an erosional remnant — located on the eastern border of the Llano Estacado within Blanco Canyon in Crosby County, Texas. With Blanco Canyon, it is the type locality of the early Pleistocene Blanco Formation of Texas and Kansas, as well as the Blancan fauna, which occurs throughout North America. Mount Blanco is a Late Blancan age site, and is associated with other Late Blancan sites from Texas such as Red Light and Hudspeth local faunas from Hudspeth County, and the Cita Canyon fauna from Randall County.
Megacamelus is an extinct genus of terrestrial herbivore in the family Camelidae, endemic to North America from the Miocene through Pliocene 10.3—4.9 mya, existing for approximately 5.4 million years.
Paleontology in Missouri refers to paleontological research occurring within or conducted by people from the U.S. state of Missouri. The geologic column of Missouri spans all of geologic history from the Precambrian to present with the exception of the Permian, Triassic, and Jurassic. Brachiopods are probably the most common fossils in Missouri.
Paleontology in Nebraska refers to paleontological research occurring within or conducted by people from the U.S. state of Nebraska. Nebraska is world-famous as a source of fossils. During the early Paleozoic, Nebraska was covered by a shallow sea that was probably home to creatures like brachiopods, corals, and trilobites. During the Carboniferous, a swampy system of river deltas expanded westward across the state. During the Permian period, the state continued to be mostly dry land. The Triassic and Jurassic are missing from the local rock record, but evidence suggests that during the Cretaceous the state was covered by the Western Interior Seaway, where ammonites, fish, sea turtles, and plesiosaurs swam. The coasts of this sea were home to flowers and dinosaurs. During the early Cenozoic, the sea withdrew and the state was home to mammals like camels and rhinoceros. Ice Age Nebraska was subject to glacial activity and home to creatures like the giant bear Arctodus, horses, mammoths, mastodon, shovel-tusked proboscideans, and Saber-toothed cats. Local Native Americans devised mythical explanations for fossils like attributing them to water monsters killed by their enemies, the thunderbirds. After formally trained scientists began investigating local fossils, major finds like the Agate Springs mammal bone beds occurred. The Pleistocene mammoths Mammuthus primigenius, Mammuthus columbi, and Mammuthus imperator are the Nebraska state fossils.
Paleontology in Idaho refers to paleontological research occurring within or conducted by people from the U.S. state of Idaho. The fossil record of Idaho spans much of the geologic column from the Precambrian onward. During the Precambrian, bacteria formed stromatolites while worms left behind trace fossils. The state was mostly covered by a shallow sea during the majority of the Paleozoic era. This sea became home to creatures like brachiopods, corals and trilobites. Idaho continued to be a largely marine environment through the Triassic and Jurassic periods of the Mesozoic era, when brachiopods, bryozoans, corals, ichthyosaurs and sharks inhabited the local waters. The eastern part of the state was dry land during the ensuing Cretaceous period when dinosaurs roamed the area and trees grew which would later form petrified wood.
Paleontology in California refers to paleontologist research occurring within or conducted by people from the U.S. state of California. California contains rocks of almost every age from the Precambrian to the Recent.