Carios

Carios
	Carios vespertilionis
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Subphylum:	Chelicerata
Class:	Arachnida
Order:	Ixodida
Family:	Argasidae
Genus:	Carios ; Latreille, 1796

Last updated October 17, 2025

Carios is a genus of ticks belonging to the family Argasidae.^[1]

The genus has cosmopolitan distribution.^[2]

Species

Carios is one of four genera in the Argasid subfamily Ornithodorinae. The genus currently contains 105 species over 7 subgenera.^[1]

Alectorobius

C. (Alectorobius) amblus Chamberlin, 1920
C. (Alectorobius) antiquus Poinar, 1995
C. (Alectorobius) aragaoi Fonseca, 1960
C. (Alectorobius) atacamensis Muñoz-Leal, Venzal & González-Acuña, 2016
C. (Alectorobius) azteci Matheson, 1935
C. (Alectorobius) brodyi Matheson, 1935
C. (Alectorobius) capensis Neumann, 1901
C. (Alectorobius) casebeeri Jones & Clifford, 1972
C. (Alectorobius) cavernicolous Dantas-Torres, Venzal & Labruna
C. (Alectorobius) cheikhi Vermeil, Marjolet & Vermeil, 1997
C. (Alectorobius) chironectes Jones & Clifford, 1972
C. (Alectorobius) clarki Jones & Clifford, 1972
C. (Alectorobius) collocaliae Hoogstraal, Kadarsan, Kaiser & Van Peenen, 1974
C. (Alectorobius) concanensis Cooley & Kohls, 1941
C. (Alectorobius) coniceps (Canestrini, 1890)
C. (Alectorobius) cyclurae de la Cruz, 1984
C. (Alectorobius) darwini Kohls, Clifford & Hoogstraal, 1969
C. (Alectorobius) denmarki Kohls, Sonenshine & Clifford, 1965
C. (Alectorobius) dugesi Mazzotti, 1943
C. (Alectorobius) dusbabeki Černý, 1967
C. (Alectorobius) dyeri Cooley & Kohls, 1940
C. (Alectorobius) echimys Kohls, Clifford & Jones, 1969
C. (Alectorobius) elongatus Kohls, Sonenshine & Clifford, 1965
C. (Alectorobius) eptesicus Kohls, Clifford & Jones, 1969
C. (Alectorobius) faccinii Barros-Battesti, Landulfo & Luz, 2015
C. (Alectorobius) fonsecai (Labruna & Venzal, 2009)
C. (Alectorobius) galapagensis Kohls, Clifford & Hoogstraal, 1969
C. (Alectorobius) guaporensis Nava, Venzal & Labruna, 2013
C. (Alectorobius) hasei (Schulze, 1935)
C. (Alectorobius) jerseyi (Klompen & Grimaldi, 2001)
C. (Alectorobius) jul Schulze, 1940
C. (Alectorobius) kelleyi Cooley & Kohls, 1941
C. (Alectorobius) knoxjonesi Jones & Clifford, 1972
C. (Alectorobius) kohlsi Guglielmone & Keirans, 2002
C. (Alectorobius) lahillei Venzal, González-Acuña & Nava, 2015
C. (Alectorobius) maritimus Vermeil & Marguet, 1967
C. (Alectorobius) marmosae Jones & Clifford, 1972
C. (Alectorobius) microlophi Venzal, Nava & González-Acuña, 2013
C. (Alectorobius) mimon Kohls, Clifford & Jones, 1969
C. (Alectorobius) muesebecki Hoogstraal, 1969
C. (Alectorobius) natalinus Černý & Dusbábek, 1967
C. (Alectorobius) peropteryx Kohls, Clifford & Jones, 1969
C. (Alectorobius) peruvianus Kohls, Clifford & Jones, 1969
C. (Alectorobius) puertoricensis Fox, 1947
C. (Alectorobius) quilinensis Venzal, Nava & Mangold, 2012
C. (Alectorobius) rietcorreai Labruna, Nava & Venzal, 2016
C. (Alectorobius) rioplatensis Venzal, Estrada-Peña & Mangold, 2008
C. (Alectorobius) rondoniensis (Labruna, Terassini, Camargo, Brandão, Ribeiro & Estrada-Peña, 2008)
C. (Alectorobius) rossi Kohls, Sonenshine & Clifford, 1965
C. (Alectorobius) rudis Karsch, 1880
C. (Alectorobius) saraivai Muñoz-Leal & Labruna, 2017
C. (Alectorobius) sawaii Kitaoka & Suzuki, 1973
C. (Alectorobius) setosus Kohls, Clifford & Jones, 1969
C. (Alectorobius) spheniscus Hoogstraal, Wassef, Hays & Keirans, 1985
C. (Alectorobius) stageri Cooley & Kohls, 1941
C. (Alectorobius) tadaridae Černý & Dusbábek, 1967
C. (Alectorobius) talaje (Guérin-Méneville, 1849)
C. (Alectorobius) tiptoni Jones & Clifford, 1972
C. (Alectorobius) tuttlei Jones & Clifford, 1972
C. (Alectorobius) xerophylus Venzal, Mangold & Nava, 2015
C. (Alectorobius) yumatensis Cooley & Kohls, 1941
C. (Alectorobius) yunkeri Keirans, Clifford & Hoogstraal, 1984

Antricola

C. (Antricola) armasi de la Cruz and Estrada-Peña, 1995
C. (Antricola) centralis de la Cruz and Estrada-Peña, 1995
C. (Antricola) cernyi de la Cruz, 1978
C. (Antricola) coprophilus McIntosh, 1935
C. (Antricola) delacruzi Estrada-Peña, Barros-Battesti and Venzal, 2004
C. (Antricola) granasi de la Cruz, 1973
C. (Antricola) guglielmonei Estrada-Peña, Barros-Battesti and Venzal, 2004
C. (Antricola) habanensis de la Cruz, 1976
C. (Antricola) hummelincki de la Cruz and Estrada-Peña, 1995
C. (Antricola) inexpectata Estrada-Peña, Barros-Battesti and Venzal, 2004
C. (Antricola) martelorum de la Cruz, 1978
C. (Antricola) mexicanus Hoffmann, 1958
C. (Antricola) naomiae de la Cruz, 1978
C. (Antricola) occidentalis de la Cruz, 1978
C. (Antricola) siboneyi de la Cruz and Estrada-Peña, 1995
C. (Antricola) silvai Černý, 1967

Carios

C. (Carios) australiensis Kohls and Hoogstraal, 1962
C. (Carios) daviesi Kaiser and Hoogstraal, 1973
C. (Carios) dewae Kaiser and Hoogstraal, 1974
C. (Carios) macrodermae Hoogstraal, Moorhouse, Wolf and Wassef, 1977
C. (Carios) pusillus Kohls, 1950
C. (Carios) quadridentatus Heath, 2012
C. (Carios) sinensis Jeu and Zhu, 1982
C. (Carios) vespertilionis (Latreille, 1796)

Notoaspis

C. (Nothoaspis) amazoniensis Nava, Venzal and Labruna, 2010
C. (Nothoaspis) reddelli Keirans and Clifford, 1975

Parantricola

C. (Parantricola) marginatus (Banks, 1910)

Reticulinasus

C. (Reticulinasus) batuensis Hirst, 1929
C. (Reticulinasus) camicasi (Sylla, Cornet and Marchand, 1997)
C. (Reticulinasus) chiropterphila Dhanda and Rajagopalan, 1971
C. (Reticulinasus) faini Hoogstraal, 1960
C. (Reticulinasus) hadiae (Klompen, Keirans and Durden, 1995)
C. (Reticulinasus) madagascariensis Hoogstraal, 1962
C. (Reticulinasus) multisetosus (Klompen, Keirans and Durden, 1995)
C. (Reticulinasus) papuensis (Klompen, Keirans and Durden, 1995)
C. (Reticulinasus) piriformis Warburton, 1918
C. (Reticulinasus) rennellensis Clifford and Sonenshine, 1962
C. (Reticulinasus) salahi Hoogstraal, 1953
C. (Reticulinasus) solomonis Dumbleton, 1959

Subparmatus

C. (Subparmatus) marinkellei Kohls, Clifford and Jones, 1969
C. (Subparmatus) mormoops Kohls, Clifford and Jones, 1969
C. (Subparmatus) viguerasi Cooley and Kohls, 1941

References

1 2 Mans, Ben J.; Featherston, Jonathan; Kvas, Marija; Pillay, Kerry-Anne; de Klerk, Daniel G.; Pienaar, Ronel; de Castro, Minique H.; Schwan, Tom G.; Lopez, Job E.; Teel, Pete; Pérez de León, Adalberto A.; Sonenshine, Daniel E.; Egekwu, Noble I.; Bakkes, Deon K.; Heyne, Heloise (1 January 2019). "Argasid and ixodid systematics: Implications for soft tick evolution and systematics, with a new argasid species list". Ticks and Tick-borne Diseases. 10 (1): 219–240. doi:10.1016/j.ttbdis.2018.09.010. ISSN 1877-959X.
↑ "Carios Latreille, 1796". www.gbif.org. Retrieved 11 May 2021.

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[:0-1] 1 2 Mans, Ben J.; Featherston, Jonathan; Kvas, Marija; Pillay, Kerry-Anne; de Klerk, Daniel G.; Pienaar, Ronel; de Castro, Minique H.; Schwan, Tom G.; Lopez, Job E.; Teel, Pete; Pérez de León, Adalberto A.; Sonenshine, Daniel E.; Egekwu, Noble I.; Bakkes, Deon K.; Heyne, Heloise (1 January 2019). "Argasid and ixodid systematics: Implications for soft tick evolution and systematics, with a new argasid species list". Ticks and Tick-borne Diseases. 10 (1): 219–240. doi:10.1016/j.ttbdis.2018.09.010. ISSN 1877-959X.

[GBIF-2] "Carios Latreille, 1796". www.gbif.org. Retrieved 11 May 2021.

[1]

[2]

Carios

Carios vespertilionis
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Subphylum:	Chelicerata
Class:	Arachnida
Order:	Ixodida
Family:	Argasidae
Genus:	Carios Latreille, 1796